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New data on Sciapodinae (Diptera: Dolichopodidae) with a revised catalogue and keys to Afrotropical species of the subfamily
by Igor Ya. Grichanov
Abstract.
Holotypes of 55 species and paratypes of additional 5 species have been examined mainly from the collections of the Royal Institute for Natural Sciences (Brussels) and Royal Museum for Central Africa (Tervuren). Descriptions of 18 new species, new records for known African species are given, 15 species are replaced, several species are excluded from the fauna of Madagascar, Congo (Kinshasa) or even Africa. Subgenus Bickeliolus Grichanov is raised to genus. The following pairs of species are synonymized: Bickeliolus haemorhoidalis (Becker) [(=B. coalescens (Parent)], Chrysosoma consentium Curran (=C. fortunatum Parent), C. katangense Curran (=C. speciosum Parent), C. mesotrichum Bezzi (=C. garambaense Vanschuytbroeck), C. singulare Parent (=C. gracile Vanschuytbroeck), Ethiosciapus bicalcaratus (Parent) [=E. setifrons (Parent)], E. finitimus (Parent) (=E. skufjini Grichanov), Plagiozopelma pallidicorne (Curran) [=P. lindneri (Vanschuytbroeck)]. A revised catalogue and keys to 12 genera and 205 Afrotropical species of Sciapodinae are also presented.
Key words: Diptera, Dolichopodidae, Sciapodinae, nomenclature, Tropical Africa.
Introduction
The last catalogue of Afrotropical dolichopodid fauna (Dyte and Smith, 1980) included about 180 species belonging to 8 genera of Sciapodinae, of which 3 genera were later synonymized, status of 1 genus was changed, and all species of the genus Sciapus were transferred to other genera. As a result, only 65 previously known Afrotropical species are placed now in the sciapodine genera sensu Dyte and Smith (1980). Bickel (1994a) recently revised the world fauna of sciapodine genera. In his big work he proposed 21 valid genera, of which 8 occur in Afrotropical Region. Grichanov (19951997) reviewed all African sciapodine genera, described two new genera and 22 new species of the subfamily from the Region. In this paper one new genus is established and one more is recorded for Africa for the first time. Now twelve genera are known from Afrotropical zoogeographical Region, of which Ethiosciapus, Bickeliolus and Gigantosciapus are endemic for Continental Africa and Madagascar, and Bickelia and Mascaromyia are confined to western Indian Ocean islands. Dytomyia was found on Madagascar and in Australia. The number of Afrotropical species belonging to sciapodine genera are as follows: Chrysosoma (67), Amblypsilopus (43), Mascaromyia (21), Plagiozopelma (17), Condylostylus (15), Gigantosciapus (13), Ethiosciapus (10), Bickeliolus (7), Mesorhaga (5), Dytomyia (3), Bickelia (3) and Sciapus (1).
In total, 205 Afrotropical species are known today. Many of those species were briefly or incompletely described. Some of them are known from females, others from males only. Authors in their descriptions often omitted important characters regarded now as having generic level value. That is why no less than 20 species are waiting to be synonymized or be declared Nomina Dubia in future revisions of type material. A few dozens of species are probably waiting to be described. The major fraction of Afrotropical species was described by O. Parent (71 species including 7 placed in synonymy), I.Ya. Grichanov (41 and 1), C.H. Curran (37 and 7), Th. Becker (21 and 2), P. Vanschuytbroeck (15 and 3) and C.G. Lamb (9).
The well-known fauna of Congo (Kinshasa) comprises 84 species; Tanzania numbers 25, South Africa 22, Uganda 20, Nigeria 19, Kenya and Cameroons 16 species each. Adjacent African islands are poorly studied. Only 29 species were recorded from Madagascar, 16 from Mauritius and 11 from Seychelles. The most interest distribution of species are as follows: Chrysosoma snelli Curran Tanzania, Kenya, Madagascar, Seychelles, Reunion, Mauritius, Rodriguez, Aldabra, Maldives, Chagos Archipelago; Mascaromyia librativertex (Lamb) Seychelles, Mauritius, Chagos Archipelago; Bickelia subparallela Grichanov Chagos Archipelago, Seychelles; Amblypsilopus munroi (Curran) South Africa, Namibia, Angola, Mozambique, Sri Lanka; Amblypsilopus pallidicornis (Grimshaw) Madagascar, Seychelles, Hawaiian Islands, Society Islands, Marquesas Islands, Guam, Belau and Taiwan.
Treating material from the collections of the Royal Institute for Natural Sciences [RINS] (Brussels), Royal Museum for Central Africa [RMCA] (Tervuren) and National Museum of Kenya [NMK] (Nairobi), I found a lot of interesting material in the subfamily Sciapodinae. Holotypes of 55 species and paratypes of additional 5 species were examined. In this paper descriptions of 18 new species, new records for known African species are given, 8 species are synonymized and 15 species are replaced, and some previous species records from Madagascar, Congo (Kinshasa) or even all Africa are shown to be incorrect. A revised catalogue and keys to all Afrotropical sciapodine species is also presented.
Holotypes and paratypes of the new species are deposited in the Royal
Institute for Natural Sciences (Brussels). Hypopygii removed from the dry
specimens are placed after alkalisation into glycerol and mounted on the
same pin in a cavity of polymer film covered with a piece of adhesive tape.
Most part of the material collected from Madagascar is kept in 70% alcohol
inside glass tubes and cans. In addition, one specimen of each species
is also placed after alkalisation into glycerol and mounted on pin in sealed
plastic container. Listing material examined, I use here slash ( / ) to
separate labels on one pin and square brackets [...] to insert my personal
remarks. Species diagnosis includes usually key characters and some important
features that were missed in original descriptions. Bibliography includes
works published after the "Catalogue of the Diptera of the Afrotropical
Region" (Dyte and Smith, 1980).
CATALOGUE OF AFROTROPICAL SPECIES OF SCIAPODINAE
TRIBE MESORHAGINI BICKEL
Genus Mesorhaga Schiener
Mesorhaga Schiener, 1868:217. Type species Mesorhaga tristis Schiener, 1868, by monotypy.
=Aptorthus Aldrich, 1893:48. Type species Aptorthus albiciliatus Aldrich, 1893, subsequently designated by Coquillett, 1910:509.
africana Curran, 1927:264 Congo (Kinshasa).
demeyeri Grichanov, sp. n. Madagascar.
mahunkai Grichanov, 1997c:45 Tanzania.
pauliani Vanschuytbroeck, 1952:137 Madagascar.
tsurikovi Grichanov, sp. n. Burundi.
TRIBE SCIAPODINI BICKEL
Genus Sciapus Zeller
=Leptopus Fallen, 1823:23. Included species Leptopus longulus Fallen, 1823 and Leptopus wiedemanni Fallen, 1823 [preoccupied by Latreille, 1809].
=Psilopus Meigen, 1824:35. Type species Dolichopus platypterus Fabricius, 1805, by subsequent designation of Westwood, 1840:134 [preoccupied by Poli, 1795].
Sciapus Zeller, 1842: 831 (new name for Psilopus Meigen)
=Stenarus Gistl, 1848:10 ([unnecessary] new name for Psilopus Meigen).
=Psilopodius Rondani, 1861:11 ([unnecessary] new name for PsilopusMeigen).
=Psilopodinus Bigot, 1889:24. Type species Dolichopus platypterus Fabricius, 1805, by original designation.
=Psilopiella Van Duzee, 1914:438. Type species Psilopiella rutila Van Duzee, 1914, by original designation.
=Agastoplax Enderlein, 1936:114. Type species Psilopus flavicinctus Loew, 1857, by monotypy.
=Dactylodiscia Enderlein, 1936:114. Type species Psilopus calceolata Loew, 1859, by original designation.
=Dactylorhipis Enderlein, 1936:114. Type species Psilopus bellus Loew, 1873, by monotypy.
=Placantichir Enderlein, 1936:114 [unavailable name, genus proposed after 1930 without designation of type species from the five included species].
=Placantichir Bickel, 1994a:88. Type species Dolichopus nervosus Lehmann, 1822, designated by Bickel, 1994a [for description to validate proposal of new genus, see generic characters cited for Placantichir Enderlein, 1936:114].
Records of Palearctic species such as S. longimanus Becker, S. adumbratus Becker and S. sylvaticus Becker from Congo (Kinshasa), Zimbabwe and Madagascar (Vanschuytbroeck, 1951, 1952, 1959) belong to undescribed species of Amblypsilopus.
endrodyi Grichanov, 1997c:47 Ghana.
Genus Dytomyia Bickel
Dytomyia Bickel, 1994a:91. Type species Sciopus sordidus Parent, 1928, by original designation.
deconinckae Grichanov, sp. n. Madagascar.
elenae Grichanov, sp. n. Madagascar.
paulyi Grichanov, sp. n. Madagascar.
Genus Bickelia Grichanov
Bickelia Grichanov, 1996a:119. Type species Bickelia subparallela Grichanov, 1996, by original designation.
guerini Parent, 1935:86 (Sciopus) Mauritius.
parallela Macquart, 1842:175 (Psilopus) Mauritius.
subparallela Grichanov, 1996a:120 Chagos Archipelago, Seychelles.
Genus Mascaromyia Bickel
Mascaromyia Bickel, 1994a:100. Type species Psilopus pollicifer Lamb, 1922:375, by original designation.
albitarsis Parent, 1935:83 (Sciopus) Mauritius.
amplicaudata Lamb, 1922:378 (Psilopus) Seychelles.
babichae Grichanov, 1996:113 Mauritius.
bickeli Grichanov, 1996:111 Mauritius.
desjardsini Macquart, 1842:175 (Psilopus) Mauritius.
duplicata Parent, 1932:230 (Sciopus) Reunion.
dytei Grichanov, 1996:116 Mauritius.
frolovi Grichanov, 1996:117 Mauritius.
grandicaudata Lamb, 1922:378 (Psilopus) Seychelles.
hutsoni Grichanov, 1996:115 Mauritius.
indistincta Lamb, 1922:376 (Psilopus) Seychelles.
kalinkini Grichanov, 1996:116 Mauritius.
leptogaster Thomson, 1869:510 (Psilopus) Mauritius.
librativertex Lamb, 1922:374 (Psilopus) Seychelles, Mauritius; Chagos Archipelago.
makhotkini Grichanov, 1996:112 Mauritius.
magnicaudata Lamb, 1922:377 (Psilopus) Seychelles.
mauritiensis Parent, 1939:270 (Sciopus) Mauritius.
pollicifer Lamb, 1922:375 (Psilopus) Seychelles.
rufiventris Macquart, 1842:174 (Psilopus) Mauritius.
shabuninae Grichanov, 1996:114 Mauritius.
vagabunda Lamb in Bezzi & Lamb, 1925:546 (Psilopus) Rodriguez.
Genus Condylostylus Bigot
Condylostylus Bigot, 1859:215. Type species Psilopus bituberculatus Macquart, 1842, by original designation.
=Dasypsilopus Bigot, 1859:215. Type species Psilopus pilipes Macquart, 1842, by original designation.
=Eurostomerus Bigot, 1859:215. Type species "Psilopus coerulus Macquart", nomen nudum (=Eurostomerus coerulus Bigot, validated by generic description).
=Oedipsilopus Bigot 1859:224. Type species Psilopus posticatus Wiedemann, 1830, by original designation.
=Tylochaetus Bigot, 1889:215. Type species Psilopus bituberculatus Macquart, 1842, by original designation.
=Laxina Curran, 1934:230. Type species Dolichopus patibulatus Fabricius, by original designation.
angustipennnis Loew, 1858:372 (1860:346) (Psilopus) South Africa.
basovi Grichanov, sp. n. Madagascar.
beckeri Speiser, 1920:218 Cameroons.
burgeoni Parent, 1935:115 Congo (Kinshasa), Tanzania, Kenya, Ruanda, Burundi.
chaineyi Grichanov, sp. n. Madagascar.
congensis Curran, 1927:263 Congo (Brazzaville), Cameroons, Congo (Kinshasa), Uganda, Tanzania, Kenya, South Africa.
=imitans Parent, 1935:117 nec Curran (misidentification).
galinae Grichanov, 1996c:218. Uganda, Congo (Kinshasa).
imitator Curran, 1924:221 Congo (Kinshasa), Kenya, Tanzania, Burundi, Malawi, Zimbabwe, Mozambique, South Africa, Angola, Namibia.
=imitans Curran, 1925:114 ([unnecessary] new name for C. imitator Curran).
kivuensis Vanschuytbroeck, 1964:136 Congo (Kinshasa).
paricoxa Parent, 1939:267 Kenya, Zimbabwe, Tanzania.
pateraeformis Becker, 1923:38 Cameroons, Nigeria, Congo (Kinshasa), Uganda, Tanzania, Kenya, South Africa, (?) Madagascar.
=alter Becker, 1923:38.
selectus Parent, 1931:43 Malawi, Congo (Kinshasa).
selitskayae Grichanov, sp. n. Congo (Kinshasa).
skufjini Grichanov, sp. n. Madagascar.
stenurus Loew, 1858:377 (1860:346) (Psilopus) South Africa, Zimbabwe.
=sicatris Curran, 1926:389.
TRIBE CHRYSOSOMATINI GUERIN-MENEVILLE
Genus Ethiosciapus Bickel
Ethiosciapus Bickel, 1994a:141. Type species Psilopus bilobatus Lamb, 1922, by original designation.
bicalcaratus Parent, 1933:37 (Sciapus) Congo (Kinshasa), Uganda, Burundi, Madagascar, Comores, St. Helena.
=setifrons Parent, 1937:14 (Sciapus), n. comb., n. syn.
bilobatus Lamb, 1922:372 (Psilopus) Seychelles.
degener Parent, 1934:119 (Condylostylus) South Africa, n. comb.
dilectus Parent, 1935:84 (Sciapus) Tanzania, Uganda.
exarmatus Parent, 1933:39 (Sciapus) Congo (Kinshasa), ?Madagascar.
finitimus Parent, 1939:262 (Chrysosoma) Congo (Kinshasa), Uganda, n. comb.
=skufjini Grichanov, 1996:226, n. syn.
flavirostris Loew, 1858:371 (Psilopus) South Africa, Mozambique, Madagascar, ?Ethiopia.
inflexus Becker, 1923:46 (Sciapus) Tanzania, Uganda, Kenya, Burundi, Congo (Kinshasa), Madagascar, South Africa, St. Helena, n. comb.
integer Becker, 1923:47 (Sciapus) Malawi.
latipes Parent, 1930:94 (Chrysosoma) Madagascar.
Genus Bickeliolus Grichanov, n. status
Bickeliolus Grichanov, 1996:224 (as subgenus). Type species Ethiosciapus maslovae Grichanov, 1996, by original designation.
alluaudi Parent, 1935:80 (Chrysosoma) Madagascar, n. comb.
haemorhoidalis Becker, 1923:46 (Sciapus) Ethiopia, Uganda, Burundi, Congo (Kinshasa), South Africa, n. comb.
=coalescens Parent, 1934:121 (Sciapus), n. comb., n. syn.
lamellatus Parent, 1935:118 (Sciapus) Congo (Kinshasa), Ruanda, Tanzania, Uganda, St. Helena, n. comb.
lasiophtalmus Lamb, 1922:371 (Psilopus) Seychelles, n. comb.
lutescens Vanschuytbroeck, 1952:139 (Sciapus) Madagascar, n. comb.
maslovae Grichanov, 1996:224 (Ethiosciapus) Angola, n. comb.
trochanteralis Curran, 1924:219 (Sciapus) South Africa, n. comb.
Genus Chrysosoma Guerin-Meneville
Subgenus Chrysosoma Guerin-Meneville
Chrysosoma Guerin-Meneville, 1831:pl.20, fig.6. Type species Chrysosoma fasciata Guerin-Meneville, 1831, by subsequent designation of Enderlein, 1912:373.
=Agonosoma Guerin-Meneville, 1838:293 ([unnecessary] replacement name for Chrysosoma Guerin-Meneville), preoccupied by Laporte, 1832.
=Margaritostylus Bigot, 1859:215. Type species Psilopus globifer Wiedemann, 1830, by original designation.
=Megistostylus Bigot, 1859:215. Type species Dolichopus crinicornis Wiedemann (as Psilopus crinicornis), by original designation.
=Mesoblepharius Bigot, 1859:215. Type species Psilopus senegalensis Macquart, 1834, by original designation.
=Oariostylus Bigot, 1889:215. Type species Psilopus tuberculicornis Macquart, 1855, by original designation.
=Eudasypus Bigot, 1889:24. Type species Psilopus senegalensis Macquart, 1834, by original designation.
=Oariopherus Bigot, 1889:24. Type species Psilopus tuberculicornis Macquart, 1855, by original designation.
=Spathiopsilopus Bigot, 1888:24. Type species Psilopus globifer Wiedemann, 1830, by original designation.
=Spathipsilopus Bigot, 1890:268. Type species Psilopus globifer Wiedemann, 1830, by subsequent designation of Dyte, 1975:213.
aequatoriale Parent, 1933:2 Congo (Kinshasa), Uganda.
aequilobatum Parent, 1933:15 Congo (Kinshasa), Congo (Brazzaville).
aestimabile Parent, 1934:18 Nigeria, Gambia, ?Madagascar.
albilimbatum Bigot, 1890:287 (Psilopus) Ivory Coast, Gabon, Ghana, Nigeria, Sierra Leone, Congo (Brazzaville), Congo (Kinshasa).
albocrinitatum Curran, 1925:109 Congo (Kinshasa), Congo (Brazzaville).
alboguttatum Parent, 1930:93 Cameroons, Burundi, Guinea.
angolense Parent, 1934:113 Angola, Congo (Kinshasa).
arduus Parent, 1936:6 (Sciapus) Congo (Kinshasa), n.comb.
asperum Parent, 1934:114 South Africa.
bacchi Dyte, 1957:37 Tanzania, Burundi.
benignum Parent, 1934:115 Nigeria.
bredoi Parent, 1933:21 Congo (Kinshasa).
carum Walker, 1849:643 (Psilopus) "Congo".
centrale Becker, 1923:25 Cameroons.
cilifemoratum Parent, 1934:115 Malawi, Congo (Kinshasa).
consentium Curran, 1925:111 Congo (Kinshasa), Congo (Brazzaville).
=fortunatum Parent, 1933:1, n. syn.
continuum Curran, 1927:255 Congo (Kinshasa), Congo (Brazzaville).
corruptor Parent, 1933:24 Congo (Kinshasa).
flexum Loew, 1858:371 (Psilopus) South Africa.
gemmeum Walker, 1849:644 (Psilopus) Nigeria, Sierra Leone, Gambia, Senegal.
gromieri Parent, 1930:92 Cameroons.
hargreavesi Curran, 1927:9 Sierra Leone.
hirsutulum Parent, 1933:27 Congo (Kinshasa).
ituriense Parent, 1933:29 Congo (Kinshasa), Malawi.
katangense Curran, 1925:107 Congo (Kinshasa), Sudan.
=speciosum Parent, 1933:32, n. syn.
kuznetzovi Grichanov, 1997:37 Zambia.
laeve Bigot, 1891:373 (Psilopodius) ?Guinea, Ivory Coast.
lavinia Curran, 1927:260 Congo (Kinshasa), Tanzania, Burundi.
leucopogon Wiedemann, 1824:40 (Dolichopus) ?Mauritius, ?Aldabra; Chagos Archipelago, Burma, Sri Lanka, Taiwan, India, Indochina, Java, Sumatra, Thailand, Queensland, New Caledonia, Tahiti.
=apicalis Wiedemann, 1830:227 (Psilopus).
=conicornis Macquart, 1846:120 (Psilopus).
=loewi Enderlein, 1912:378.
liberia Curran, 1929:4 Liberia, Congo (Kinshasa).
marginatum Becker, 1923:29 Sierra Leone.
mesotrichum Bezzi, 1908:380 (Psilopus) Congo (Kinshasa), Sierra
Leone, Uganda.
=senegalense Becker, 1923:20,33 nec Macquart (misidentification).
=senegalense Curran, 1925:107 nec Macquart (misidentification).
=garambaense Vanschuytbroeck, 1959:37, n.syn.
micantifrons Speiser, 1910:108 (Agonosoma) Tanzania.
minusculum Becker, 1923:29 Cameroons, Equatorial Guinea, Congo (Kinshasa), ?Sierra Leone.
=ostentatum Becker, 1923:30.
norma Curran, 1927:257 Congo (Kinshasa).
pauperculum Parent, 1933:30 Congo (Kinshasa).
petersi Dyte, 1957:37 Tanzania.
pomeroyi Curran, 1927:7 Nigeria, Cameroons.
praecipuum Parent, 1936:319 Congo (Kinshasa).
praelatum Becker, 1923:31 Equatorial Guinea, Tanzania, Congo (Kinshasa), Malawi.
pseudorepertum Grichanov, sp. n. Congo (Kinshasa).
repertum Becker, 1923:31 Equatorial Guinea, Nigeria.
schoutedeni Curran, 1927:258 Congo (Brazzaville), Congo (Kinshasa), Tanzania, Angola, Zambia.
=fortunatum Grichanov, 1997 nec PARENT (misidentification).
senegalense Macquart, 1834:450 (Psilopus) Congo (Brazzaville), Congo (Kinshasa), Gabon, Senegal, Sierra Leone, Nigeria.
=smaragdinum Walker, 1849:642 (Psilopus).
=saphirum Bigot, 1858:362 (Psilopus).
=mixtum Curran, 1927:3.
singulare Parent, 1933:3 Congo (Kinshasa).
=gracile Vanschuytbroeck, 1959:29, n.syn.
snelli Curran, 1927:5 Tanzania, Kenya, Madagascar, Seychelles, Reunion, Rodriguez; Maldives, Chagos Archipelago.
=leucopogon, authors, nec Wiedemann.
stolyarovi Grichanov, sp. n. Burundi.
stubbsi Grichanov, 1997:35 Kenya, Uganda.
tarsiciliatum Parent, 1930:91 Cameroons, Sierra Leone, Congo (Kinshasa), Gabon.
tanasijtshuki Grichanov, 1997:33 Kenya, Congo (Kinshasa).
tenuipenne Curran, 1927:254 Senegal, Nigeria, Congo (Brazzaville), Congo (Kinshasa), Uganda.
tractatum Becker, 1923:35 Togo, Nigeria, Ghana.
=biciliatum Parent, 1931:45.
tricrinitum Parent, 1933:33 Congo (Kinshasa), Burundi, Mozambique, Malawi, Sierra Leone, Nigeria, Tanzania, South Africa.
=flexum Curran, 1926:384 nec Loew (misidentification).
trigemmans Walker, 1849:650 (Psilopus) no locality.
triumphator Parent, 1933:33 Congo (Kinshasa).
ungulatum Parent, 1941:207 Principe.
varivittatum Curran, 1925:112 Congo (Kinshasa).
vividum Becker, 1923:36 Equatorial Guinea, Congo (Kinshasa), Cameroons.
=crinipes Parent, 1933:25.
woodi Parent, 1935:82 Zambia.
zaitzevi Grichanov, 1997:38 Zambia.
zephyrum Bigot, 1858:361 (Psilopus) Gabon.
zinovjevi Grichanov, 1997:31 Nigeria.
Subgenus Kalocheta Becker
Kalocheta Becker, 1923:41 (as genus). Type-species: Kalocheta passiva Becker, 1923, by monotypy.
collarti Parent, 1933:35 (Kalocheta) Congo (Kinshasa), Tanzania, Kenya, Uganda.
cucana Negrobov et Kulibali, 1983:1121 (Kalochaeta) Congo (Kinshasa), Uganda.
neoliberia Bickel, 1994a:212 Liberia.
=liberia Curran, 1929:5 (Kalocheta) nec Curran, 1929:4 (preoccupied).
passiva Becker, 1923:42 (Kalocheta) Cameroons, ?Congo (Brazzaville), ?Congo (Kinshasa).
villiersi Vanschuytbroeck, 1970:267 (Kalocheta) Congo (Brazzaville).
Genus Gigantosciapus Grichanov
Gigantosciapus Grichanov, 1997:79. Type-species: Gigantosciapus oldroydi Grichanov, 1997: original designation.
africanus Parent, 1933:16 (Chrysosoma) Congo (Kinshasa).
anomalipes Parent, 1935:112 (Chrysosoma) Congo (Kinshasa), Ghana.
decellei Vanschuytbroeck, 1966:202 (Megistostylus) Ivory Coast.
francoisi Grichanov, sp. n. Congo (Kinshasa).
gemmarius Walker, 1849:645 (Psilopus) Sierra Leone, Congo (Brazzaville), Ghana, Congo (Kinshasa), Ivory Coast, Liberia, Cameroons.
=fulvicinctus Bigot, 1891:372 (Psilopodius).
inversus Curran, 1927:249 (Chrysosoma) Sierra Leone, Liberia.
kamerunensis Becker, 1923:27 (Chrysosoma) Cameroons, Guinea, Congo (Kinshasa), ?Madagascar.
meyeri Vanschuytbroeck, 1962:353 (Megistostylus) Nigeria.
nataliae Grichanov, sp. n. Congo (Kinshasa).
oldroydi Grichanov, 1997:80 Cameroons.
pseudogemmarius Parent, 1934:118 (Chrysosoma) Congo (Brazzaville).
saegeri Vanschuytbroeck, 1959:9 (Megistostylus) Congo (Kinshasa).
tuberculatus Curran, 1927:246 (Chrysosoma) Ghana, Ivory Coast.
Genus Plagiozopelma Enderlein
Plagiozopelma Enderlein, 1912:367. Type species Plagiozopelma spengeli Enderlein, 1912, by original designation [=Psilopus appendiculatus Bigot, 1890].
angulitarse Parent, 1933:18 (Chrysosoma) Congo (Kinshasa).
bequaerti Curran, 1926:2 (Chrysosoma) Congo (Kinshasa), Uganda.
capilliferum Parent, 1933:22 (Chrysosoma) Congo (Kinshasa).
collarti Curran, 1927:249 (Chrysosoma) Congo (Brazzaville), Malawi, Uganda, Congo (Kinshasa).
conjectum Parent, 1934:116 (Chrysosoma) Ghana.
daveyi Parent, 1939:261 (Chrysosoma) Malawi.
du Curran, 1929:2 (Chrysosoma) Liberia, Congo (Kinshasa).
ghesquieri Parent, 1936:2 (Chrysosoma) Congo (Kinshasa).
grahami Parent, 1939:264 (Chrysosoma) Ghana.
inops Parent, 1929:202 (Chrysosoma) Liberia, Benin, Tanzania, Congo (Kinshasa).
nalense Curran, 1926:6 (Chrysosoma) Congo (Kinshasa), Nigeria, Tanzania.
njalense Parent, 1934:118 (Chrysosoma) Sierra Leone.
pallidicorne Curran, 1927:252 (Chrysosoma) Kenya, Tanzania, Congo (Kinshasa), n. comb.
=lindneri Vanschuytbroeck, 1964:3 (Megistostylus), n. syn.
=puma Dyte & Smith, 1980:446 (Chrysosoma) ([unnecessary] new name for Chrysosoma pallidicorne Curran), n. comb.
piliseta Parent, 1936:4 (Chrysosoma) Congo (Kinshasa).
ramiseta Parent, 1939:266 (Chrysosoma) Sierra Leone, Zimbabwe, Congo (Kinshasa).
tritiseta Parent, 1929:271 (Chrysosoma) Cameroons, Ghana, Nigeria, Congo (Kinshasa).
vagator Becker, 1923:36 (Chrysosoma) Togo, Congo (Kinshasa), Equatorial Guinea.
Genus Amblypsilopus Bigot
Amblypsilopus Bigot, 1889:24. Type species Psilopus pittacinus Loew, 1861 (as pitacinus Fabricius), by original designation.
=Gnamptopsilopus Aldrich, 1893:48. Type species Psilopus scintillans Loew, designated by Coquillett, 1910:547.
=Leptorhethum Aldrich, 1893:50. Type species Leptorhethum angustatum Aldrich, 1893, by monotypy.
=Sciopolina Curran, 1924:216. Type species Sciopolina fasciatum Curran, 1924, by monotypy.
=Australiola Parent, 1932:127. Type species Australiola tonnoiri Parent, 1932 (=Sciapus zonatus Parent, 1932), by original designation.
=Labeneura Parent, 1937:126 (as subgenus of Sciapus). Type species Labeneura barbipalpis Parent, 1937 (=Sciapus lenga Curran, 1929), by monotypy.
aenescens Vanschuytbroeck, 1952:138 (Sciapus) Madagascar.
auratus Curran, 1924:217 (Chrysosoma) South Africa, Zimbabwe, Zambia, Angola, Congo (Kinshasa), Tanzania, Nigeria, Guinea.
barkalovi Grichanov, sp. n. Congo (Kinshasa).
basilewskyi Vanschuytbroeck, 1960:319 (Sciapus) Tanzania, Kenya, Uganda.
bevisi Curran, 1927:11 (Sciapus) South Africa.
bipectinatus Parent, 1934:120 (Sciapus) Kenya.
bonniae Irwin, 1974:245 (Sciopolina) South Africa.
bruneli Grichanov, sp. n. Madagascar.
cilifrons Parent, 1937:126 (Chrysosoma) Nigeria, Togo, Congo (Kinshasa), Kenya, Madagascar.
cuthbertsoni Parent, 1937:129 (Sciapus) Zimbabwe, Burundi.
dallastai Grichanov, sp. n. Madagascar.
disjunctus Parent, 1936:1 (Chrysosoma) Congo (Kinshasa), Nigeria, Burundi.
fasciatus Curran, 1924:216 (Sciopolina) South Africa.
=palliatus Curran, 1927:12 (Sciapus).
flabellifer Becker, 1923:45 (Sciapus) Madagascar, ?Congo (Kinshasa).
flavicollis Becker, 1923:40 (Leptorhethum) Cameroons, Equatorial Guinea.
flavus Vanschuytbroeck, 1962:353 (Megistostylus) Madagascar, n.comb.
gorodkovi Grichanov, 1996:291 Tanzania.
grootaerti Grichanov, sp. n. Madagascar.
kraussi Grichanov, sp. n. Madagascar.
lenga Curran, 1929:1 (Sciapus) Liberia, Sierra Leone, Zimbabwe.
=barbipalpis Parent, 1937:128 (Labeneura, as subgenus of Sciapus).
longifilis Becker, 1923:28 (Chrysosoma) Tanzania, Kenya, Uganda, Congo (Kinshasa), St. Helena.
macularivenus Irwin, 1974:251 (Sciopolina) South Africa.
madagascarensis Vanschuytbroeck, 1952:89 (Chrysosoma) Madagascar.
miserus Parent, 1935:81 (Chrysosoma) Mozambique, Zimbabwe.
munroi Curran, 1924:218 (Chrysosoma) South Africa, Namibia, Angola, Mozambique; Sri Lanka.
=ernestus Curran, 1924:218 (Chrysosoma).
nanus Parent, 1929:243 (Sciapus) Senegal, Congo (Kinshasa).
nartshukae Grichanov, 1996:290 Angola.
nubilis Parent, 1935:87 (Sciapus) Madagascar.
pallidicornis Grimshaw, 1901:12 (Gnamptopsilopus) Madagascar, Seychelles; Hawaiian Islands, Society Islands, Marquesas Islands, Guam, Belau, Taiwan.
=fulgidipenne Enderlein, 1912:377 (Chrysosoma).
parilis Parent, 1931:44 (Chrysosoma) South Africa, Namibia, Zimbabwe, Tanzania, Congo (Kinshasa), Nigeria.
pernigrus Becker, 1923:30 (Chrysosoma) Malawi, Congo (Kinshasa).
rectangularis Parent, 1937:13 (Sciapus) Congo (Kinshasa), ?Madagascar.
retrovenus Irwin, 1974:242 (Sciopolina) South Africa.
rosaceus Wiedemann, 1824:40 (Dolichopus) South Africa.
=peringueyi Curran, 1926:2 (Sciapus).
signatus Becker, 1923:35 (Chrysosoma) Malawi, Congo (Kinshasa).
simplex de Meijere, 1910:99 (Agonosoma) ?Seycheles; Java.
steelei Grichanov, 1996:289 Kenya.
stuckenbergi Vanschuytbroeck, 1957:3 (Sciapus) Madagascar.
stuckenbergorum Irwin, 1974:236 (Sciopolina) South Africa.
subfascipennis Curran, 1926:386 (Sciapus) Uganda, Congo (Kinshasa), St. Helena, ?Madagascar.
sudanensis Parent, 1939:271 (Sciapus) Sudan, Congo (Kinshasa).
tenuicauda Parent, 1936:7 (Sciapus) Congo (Kinshasa).
tropicalis Parent, 1933:40 (Sciapus) Congo (Kinshasa).
REVIEW OF TYPE MATERIAL EXAMINED, DESCRIPTIONS AND NEW RECORDS
1. Mesorhaga africana Curran
Type material examined. Holotypus, ?[red label] / Musee du Congo, Bas-Uele: Buta, Congo, Jan. 2331.1925, Dr. H. Schouteden / Type Mesorhagaafricana Curran.
Diagnosis. M. africana differs from other Afrotropical species of the genus in smaller size (2.25 mm) and, in addition, right-angular, rather than gentle, bend of M1+2 vein.
Distribution. Congo (Kinshasa).
2. Mesorhaga demeyeri sp.n. (Fig. 1)
Holotype. ? in glycerol. Madagascar: Ambatondvaraha, 26.II.1991, A. Pauly col. [RINS].
Paratypes [all in alcohol]. 3 ?? with the same label as holotype. 3 ??, Madagascar: Foulpointe, forét, lagune, X.1993, A. Pauly col. P.M. [RINS].
Description. Frons and face metallic dark-green. 2 short front vertical seta bends forward on each side; ocellar tubercle with 2 pairs of strong bristles and 1 or 2 pairs of weak posterior setae; 2 long postvertical setae positioned as a linear continuation of the postocular setal row. Ventral postcranium covered with irregular white hairs. Face narrowed, approximately 1.5 times as high as wide under antennae. Proboscis and palpus black-brown, with short light hairs; palpus with 1 black seta in addition. Antennae black. Pedicel with ring of short setae; 1 or 2 ventral setae slightly longer than pedicel. First flagellomere rounded, as long as high, with very short hairs. Arista apicodorsal. Length ratio of scape to pedicel to of first flagellomere to arista, 5 : 5 : 6 : 38.
Mesonotum and scutellum metallic green. Pleura bronze-black. 5 strong dorsocentral setae, 3 long acrostichals, a pair of strong and pair of fine scutellar setae.
Legs mostly black-brown; fore knee, tibia and basitarsus yellow; middle knee yellow-brown. Fore and middle coxae with numerous dark cilia anteriorly; hind coxa with external seta. All femora with double ventral row of black cilia, approximately equal in length to femora diameter. Fore tibia bare; tarsus simple. Length ratio of fore femora to tibia to tarsus (segments from first to fifth), 55 : 62 : 26 : 11 : 8 : 5 : 7. Middle tibia with 1 very short dorsal and 3 or 4 apical setae, tarsus simple. Length ratio of middle tibia to tarsus (segments from first to fifth), 80 : 46 : 17 : 11 : 7 : 7. Hind tibia with several inconspicuous dorsal and 2 or 3 apical setae. Last tarsomere of hind tarsus slightly flattened. Length ratio of hind tibia to tarsus (segments from first to fifth), 90 : 37 : 29 : 18 : 10 : 8.
Wings brownish in anterior half, densely brown at base; veins brown. R1 reaching mid-wing. R2+3 straight. R4+5 gently curved to M1 at apex. M1 with two right-angular bends. Ratio of parts of costa between R2+3 and R4+5 to those between R4+5 and M1, 50 : 8. M2 absent. Crossvein m-cu straight. Ratio of crossvein m-cu to apical part of M1+2 (up to curvation) to apical part of CuA1, 31 : 40 : 32. Alula with somewhat longer than usually, flattened cilia. Fold-like anal vein and anal lobe present. Lower calypter brownish, with fan of black setae. Halter brown.
Abdomen bronze-black, with short, though laterally long, black hairs. First tergite with medial triangular excavation and several long black distolateral bristles. Sixth and seventh segments shortened. Hypopygium black-brown, epandrium elongate. Cercus mostly brown, digitiform, slightly curved ventrad, with short black dorsolateral setae, several subapical ventral setae, apicodorsal fringe of short curved setae and transverse inner row of several short subapical spines. Surstylus bi- or trilobate, with several dorsal and lateral setae. Epandrial lobe long, flattened, bearing one strong seta in the middle and short seta at apex.
Female. Similar to male except lacking male secondary sexual characters, otherwise as follows. Middle tibia and basitarsus entirely yellow-brownish; wing only slightly darkened.
Length: body 3.8 mm; antenna 0.8 mm; wing-length 3.6 mm; wing-width 1.3 mm.
Distribution: Madagascar.
Etymology. The species is named for Belgian entomologist Marc De Meyer.
Diagnosis. The new species is related to M. pauliani, differing in partly yellow legs, white postocular setae, different ratio of wing veins m-cu to CuA1 and hypopygium morphology.
3. Mesorhaga pauliani Vanschuytbroeck (Fig. 2)
Material examined. ? in glycerol, Madagascar: Tam, Morarano-chrome, 1630.IX.1991, A. Pauly, forét, 25 km W. ? in alcohol, Madagascar: Tam, Morarano-chrome, 115.IX.1991, A. Pauly, forét, 25 km W. 10 ??, 7 ?? in alcohol, Madagascar: Fia Ranomafana, 19.I.1992, A. Pauly, forét [RINS].
Description. Male. Frons and face metallic green. A strong front vertical seta bends forward; ocellar tubercle with a pair of strong bristles; 5 or 6 long postvertical setae positioned as a linear continuation of the postocular setal row. Lower postocular setae black. Face narrowed, approximately 1.5 times as high as wide under antennae. Proboscis and palpus black-brown, with short light hairs; palpus supplemented with one black seta. Antennae black. Pedicel with short dorsal and 2 very long ventral setae, 4 times longer than pedicel. First flagellomere rounded, approximately as long as high. Arista dorsal. Length ratio of scape to pedicel to first flagellomere to arista, 6 : 5 : 6 : 40.
Mesonotum and scutellum metallic green-black. Pleura bronze-black. 5 strong dorsocentral setae, 3 long acrostichals, a pair of strong and pair of fine scutellar setae.
Legs black-brown, fore tibia and basitarsus somewhat lighter. Fore and middle coxae with black setae anteriorly; hind coxa with strong external seta at base. All femora with double ventral row of black setae, longer than femora diameter. Last tarsomere of all tarsi slightly flattened. Fore tibia with 2 posterodorsal seta. Length ratio of fore tibia to tarsus (segments from first to fifth), 57 : 30 : 10 : 7 : 5 : 6. Middle tibia with 2 anterodorsal, 1 posterodorsal setae. Length ratio of middle tibia to tarsus (segments from first to fifth), 74 : 44 : 15 : 12 : 6 : 6. Hind tibia with 4 dorsal setae. Length ratio of hind tibia to tarsus (segments from first to fifth), 85 : 33 : 26 : 16 : 9 : 8.
Wings evenly darkened in anterior half; veins brown. R1 nearly reaching mid-wing. R2+3 straight. R4+5 gently curved to M1 at apex. M1 with double, nearly right-angular bend. Ratio of parts of costa between R2+3 and R4+5 to those between R4+5 and M1, 34 : 6. M2 absent. Crossvein m-cu nearly straight. Ratio of crossvein m-cu to apical part of M1+2 (section from m-cu to curvation) to apical part of CuA1, 23 : 33 : 34. Fold-like anal vein and anal lobe present. Lower calypter brown, with fan of long black setae. Halter brown.
Abdomen bronze-black, with short, though laterally long, black hairs. First tergite without membranous excavation, with several long black distolateral bristles. Sixth and seventh segments shortened. Hypopygium black, epandrium elongate. Cercus mostly brown, dorsolaterally with several black setae in basal third, apically trilobate; the widest ventral lobe bearing 4 short thick dorsal setae and thin ventral apophysis. Surstylus broad, with four distal setae. Epandrial lobe long, flattened, bearing 2 subapical setae.
Female. Similar to male except lacking male secondary sexual characters. Wing only slightly darkened, almost hyaline.
Length: body 3.3 mm; antenna 0.9 mm; wing-length 3.0 mm; wing-width 1.1 mm.
Distribution: Madagascar.
Diagnosis. M. pauliani differs from other Afrotropical species of the genus in entirely black-brown leg. 2nd to 5th segments of fore tarsus shortened. M1+2 with two nearly right-angular bends; m-cu 2/3 as long as middle section of M1+2 (from m-cu to curvation) or CuA1.
4. Mesorhaga tsurikovi sp.n. (Fig. 3)
Holotype. ?, Urundi: Bururi, alt. 1950, 8.I.1949, F.J. Francois / R.I.Sc.N.B. I.G. 24.452. [RINS]
Paratypes. 2 ??, Urundi: Bururi, alt. 2000, 10.X.1948, F. Francois / R.I.Sc.N.B. I.G. 24.452. [RINS]
Description. Frons and face metallic green, mostly grey pollinose. A hairlike front vertical seta bends forward; ocellar tubercle with a pair of strong bristles and a pair of weak posterior hairs; 3 long postvertical setae are positioned as a linear continuation of the postocular setal row. Ventral postcranium covered with irregular white hairs. Face narrowed, 1.6 times as high as wide under antennae; clypeus separated from margins of eyes, 1/3 as high as epistome. Proboscis and palpus black-brown, with short light hairs. Antennae black. Pedicel with short dorsal and ventral bristles. First flagellomere transverse-oval, shorter than high, with very short hairs. Arista apicodorsal, microscopically haired. Length ratio of scape to pedicel to first flagellomere to arista, 5 : 5 : 5 : 7 : 65.
Mesonotum and scutellum metallic green, slightly brownish pollinose. Pleura bronze-black, densely grey pollinose. 5 strong dorsocentral setae, 3 long acrostichals, a pair of strong and pair of fine scutellar setae.
Legs mostly dark-yellow; fore coxa yellow-brownish, blackish at base; middle and hind coxae black; anterior four femora brown at base; hind femora brown at base and apex; hind tibia black at extreme apex; basitarsi light-brown in basal half; tarsi otherwise black. Fore and middle coxae with numerous dark cilia anteriorly; hind coxa with fine external seta. Fore femora with posterior hairs, longer than femora diameter; fore tibia bare; tarsus simple. Length ratio of fore coxa to femora to tibia to tarsus (segments from first to fifth), 55 : 90 : 99 : 60 : 25 : 17 : 8 : 9. Middle femora with fine brownish posteroventral hairs, as long as femora diameter. Middle tibia with 2 or 3 apical setae, tarsus simple. Length ratio of middle coxa to femora to tibia to tarsus (segments from first to fifth), 35 : 100 : 136 : 90 : 34 : 23 : 12 : 11. Hind femora with posteroventral hairs, half as long as femora diameter; tibia with 2 or 3 apical setae. Last tarsomere of hind tarsus slightly flattened. Length ratio of hind coxa to femora to tibia to tarsus (segments from first to fifth), 25 : 135 : 167 : 60 : 52 : 33 : 19 : 11.
Wings mostly hyaline, brownish in anterior half; veins brown. R1 nearly reaching mid-wing. R2+3 straight. R4+5 gently curved to M1 at apex. M1 with strong, 60º bend, than gently curved apicad. Ratio of parts of costa between R2+3 and R4+5 to those between R4+5 and M1, 6 : 1. M2 absent. Crossvein m-cu straight. Ratio of crossvein m-cu to apical part of M1+2 (up to curvation) to apical part of CuA1, 38 : 70 : 50. Fold-like anal vein and anal lobe present. Lower calypter brown, with black setae. Halter black-brown.
Abdomen bronze-black, with short, though laterally long, black hairs. First tergite with narrow membranous excavation and a few long black distolateral bristles. Fifth tergite ventrally swollen, with numerous long black setae; sixth and seventh segments shortened; first to sixth segments combined 2.5 times longer than mesonotum. Hypopygium black, epandrium elongate. Cercus mostly brown, strongly curved, dorsolaterally with several long black setae in the middle, apically with 2 lobes; broad ventral lobe without thick setae, bearing soft thin appendix on apex; dorsal lobe half as long as ventral, pointed, arising from rosette of short spinules. Surstylus elongate, with four apical lobes, each of them bearing 1 or 2 setae. Epandrial lobe long, flattened, fine at apex and bearing 2 setae in apical third.
Female unknown.
Length: body 4.7 5.3 mm; antenna 1.2 mm; wing-length 5.8 mm; wing-width 1.6 mm.
Distribution: Burundi.
Etymology. The species is named for Russian dipterologist Mikhail Tsurikov.
Diagnosis. The new species can be separated from M. mahunkai in larger size, another ratio of middle tarsomeres and CuA1/m-cu, mostly yellow femora and morphology of cercus. It cannot be associated with M. pauliani and female of M. africana having smaller size (2.253.3 mm) and right-angular, rather than gentle, bend of M1+2 vein.
5. Sciapus longimanus Becker
Remark. S. longimanus determined by P. Vanschuytbroeck (female examined, RINS) to be from Madagascar belongs to undescribed species of Amblypsilopus and should be excluded from the fauna of the island. Four females and male (examined, RINS) from Congo (Kinshasa) labelled by the same author as S. longimanus belongs to undescribed or indeterminable species of Amblypsilopus and Condylostylus. So, the species does not occur in the Afrotropical region.
Distribution. Palearctic region.
6. Dytomyia deconinckae sp.n. (Figs. 4, 5)
Holotype. ?, Fort Dauphin, R.P. / Institut Scientifique Madagascar [RINS].
Paratypes. 5 ?? with the same labels.
Description. Frons broad, shining metallic blue-green, slightly pollinose. A weak long front vertical bristle bending forward, a strong long postvertical one positioned as a linear continuation of the postocular setal row. Ventral postcranium covered with dense irregular white hairs. Face blue-green, white pollinose, slightly convex; clypeus separated from eyes, not coming down eyes, widely rounded at apex; face slightly narrowed, 1.4 times as high as wide under antennae. Palpi and proboscis yellow, with light hairs, palpus also with two black bristles. Antenna mostly yellow, with scape and 1st flagellomere darkened dorsally, in total 1.2 times as long as height of head. Scape slightly swollen; pedicel with a ring of short setae and one dorsal bristle equal in length to first flagellomere. The latter rounded, slightly longer than high, with short yellow hairs. Arista apicodorsal, bare. Length ratio of scape to pedicel to first flagellomere to arista, 4 : 4 : 7 : 75.
Mesonotum and scutellum shining blue-green. Pleura bronze-green, densely grey pollinose. 4 dorsocentral setae with anterior one hairlike; 1 or 2 pairs of long acrostichals. Scutellum with two strong setae.
Legs including trochanters yellow. Middle and hind coxae brown-black, apical segments of tarsi brown. Fore coxa yellow, from the front with long white hairs and 3 or 4 long yellow apical setae. Middle and hind coxae from the outside with a few yellow hairs, hind coxa also with 3 or 4 fine long yellow external setae in basal half. Fore femora with 3 or 4 long posteroventral setae in the middle, the longest is half as long as femora length; with several anteroventral setae, the longest is longer than femora diameter. Fore tibia with 1 posteroventral seta at basal fourth and microscopic semi-erect setulae along entire length. Fore basitarsus slightly broadened, ventrally flattened, with pale pile; other tarsomeres simple. Length ratio of fore coxa to femora to tibia to tarsus (segments from first to fifth), 52 : 83 : 63 : 52 : 24 : 15 : 8 : 7. Middle femora bare. Middle tibia with 1 anterodorsal, 1 anterior, 1 posterodorsal and 2 or 3 apical setae; all setae short. Length ratio of middle coxa to femora to tibia to tarsus (segments from first to fifth), 28 : 80 : 101 : 80 : 32 : 18 : 8 : 8. Hind femora bare. Hind tibia with several short dorsal and posterodorsal, 2 or 3 apical setae. Length ratio of hind coxa to femora to tibia to tarsus (segments from first to fifth), 25 : 100 : 138 : 61 : 32 : 20 : 12 : 10.
Wings hyaline, veins brown. R1 reaching 0.4 of wing length. R4+5 gently curved to M1 at apex. M1 gently curved and forming the right angle with M1+2. Ratio of parts of costa between R2+3 and R4+5 to those between R4+5 and M1, 16 : 4. M2 present as short stub vein and faint fold on membrane. Crossvein m-cu almost straight. Ratio of crossvein m-cu to apical part of M1+2 (fork-handle) to apical part of CuA, 31 : 38 : 19. Anal vein and lobe present. Anal angle acute. Lower calypter yellow, with brown edging and fine light hairs. Halters yellow, halter stem thin, nearly twice as long as knob, with short row of dark setulae in front of knob.
Abdomen shining blue-green, with copper reflection in places, weakly white pollinose, with short black hairs and long black marginal bristles. Base of segments mat-black; first and second segments with short white hairs; unmodified segments combined nearly twice as long as mesonotum; seventh segment invisible; 8th segment positioned symmetrically, covering epandrium basodorsally. Hypopygium brown. Surstylus brown, with two thin mid-dorsal apophysis. Epandrial lobe distinct, with 2 setae. and having several long setae. Cercus yellow, with light hairs and wide articulated apical lobe. Basoventral lobes of both cerci forming capitulate "Organ X" with longitudinal suture, fused along entire length, long, with 2 short ventral setae.
Female. Unknown.
Length: body 3.6 mm; antenna 1.05 mm; postabdomen 0.5 mm; wing-length 3.6 mm; wing-width 1.1 mm.
Distribution. Madagascar.
Etymology. The species is named for Belgian dipterologist Dr. E. De Coninck.
Diagnosis. Five species of the genus were earlier known from Australia (Bickel, 1994a). D. deconinckae has some similarities with Australian D. flaviseta Bickel, strongly differing from the latter in black setae on head and thorax, long posteroventral seta at base of fore tibia, more complex hypopygium morphology and many other characters.
7. Dytomyia elenae sp.n. (Fig. 6)
Holotype. ? in glycerol. Madagascar: Foulpointe, X.1993, A. Pauly col., forét de lagune, P.M. [RINS].
Paratype. ? in alcohol, Madagascar: Tam, Foulpointe, X.1993, A. Pauly col. [RINS].
Description. Frons broad, metallic blue-green. A strong front vertical bristle bends forward, postvertical one is positioned as a linear continuation of the postocular setal row. Ventral postcranium covered with dense irregular white hairs. Face metallic green, clypeus separated from eyes; face slightly narrowed, 1.4 times as high as wide under antennae. Palpi and proboscis orange, with light hairs, palpus with 2 black bristles. Antennae mostly blackish-brown, longer than height of head. Pedicel with ring of short setae. First flagellomere ovate, a little longer than high, with short hairs. Arista dorsal to apicodorsal, bare and simple. Length ratio of scape to pedicel to first flagellomere to arista, 5 : 5 : 7 : 65.
Mesonotum and scutellum metallic blue-green. Pleura bronze-green. 3 strong and 1 or 2 short anterior dorsocentral setae; 3 pairs of short acrostichals, restricted to anterior third of mesonotum. Scutellum with two strong setae.
Legs including trochanters and fore coxa yellow. Middle and hind coxae bronze-black, apical segments of tarsi brown. Fore coxae from the front with white hairs and several yellow setae. Middle and hind coxae with sparse yellow cilia. All femora practically bare. Fore tibia with long posteroventral seta just before the middle, nearly half as long as tibia length. Fore basitarsus ventrally flattened, with dense pale pile. Length ratio of fore coxa to femora to tibia to tarsus (segments from first to fifth), 45 : 75 : 72 : 42 : 40 : 27 : 15 : 8. Middle tibia with 2 anterodorsal and 2 or 3 apical setae. Length ratio of middle coxa to femora to tibia to tarsus (segments from first to fifth), 34 : 77 : 112 : 85 : 34 : 23 : 11 : 8. Hind tibia with weak setae. Length ratio of hind coxa to femora to tibia to tarsus (segments from first to fifth), 25 : 95 : 143 : 80 : 42 : 25 : 12 : 8.
Wings hyaline, veins brown. R1 2/5 as long as wing. R2+3 almost parallel to costa. R4+5 gently curved to M1 in apical fifth. Ratio of parts of costa between R2+3 and R4+5 to those between R4+5 and M1, 16 : 4. M1 widely arcuate. M1+2 and M1 forming right angle. M2 present as short stub vein and faint fold on membrane. Crossvein m-cu straight, oblique. Ratio of crossvein m-cu to apical part of M1+2 (fork-handle) to apical part of CuA, 30 : 35 : 15. Anal vein fold-like, anal lobe developed. Anal angle acute. Alula small but distinct. Lower calypter yellow, with brown edging and dark hairs. Halters yellow, halter stem as long as knob.
Abdomen metallic dark-green, with short black hairs and long black marginal setae. Sternite with short hairs. Hypopygium green-black. 8th segment positioned symmetrically, as a continuation of 7th tergite, covering epandrium basodorsally and having several long setae. Cercus yellow at base, with two brown narrow lobes in apical half and small middorsal process; basoventral lobes of both cerci long, bare, fused in basal 2/3.
Female. Unknown.
Length: body 3.4 mm; antenna 1.2 mm; wing-length 3.4 mm; wing-width 1.1 mm.
Distribution. Madagascar.
Diagnosis. B. elenae is related to B. paulyi (see diagnosis of this species), differing in posteroventral seta positioned just before the middle of fore tibia and narrow cercal lobes having shorter setae.
8. Dytomyia paulyi sp.n. (Fig. 7)
Holotype. ? in glycerol. Madagascar: Foulpointe, forét, lagune, X.1993, A. Pauly col., , P.M. [RINS].
Paratypes [in alcohol]. 7 ??, Madagascar: Foulpointe, 2.XI.1991, A. Pauly., plage, bac jaune. 5 ??, Madagascar: Foulpointe, XI.1995, A. Pauly. 4 ??, Madagascar: Foulpointe, XI.1995, A. Pauly rec., plage, bac jaune. [RINS].
Description. Frons broad, metallic blue-green. A strong front vertical bristle bends forward, postvertical one is positioned as a linear continuation of the postocular setal row. Ventral postcranium covered with dense irregular white hairs. Face metallic green, clypeus separated from eyes; face slightly narrowed, slightly higher than wide under antennae. Palpi and proboscis orange, palpus with light hairs and 2 black bristles. Antennae mostly black, as long as height of head. Scape yellow, brownish dorsally. Pedicel with ring of short setae; the longest dorsal seta longer than pedicel. First flagellomere rounded, as long as high, with short hairs. Arista subapical, bare and simple. Length ratio of scape to pedicel to first flagellomere to arista, 5 : 5 : 5 : 70.
Mesonotum and scutellum metallic blue-green. Pleura bronze-green. Three strong posterior and one short anterior dorsocentral setae; one pair of long and 1 or 2 pairs of microscopic acrostichals, restricted to anterior third of mesonotum. Scutellum with two strong setae.
Legs including anterior four trochanters yellow. Middle and hind coxae bronze-black, hind trochanter and apical segments of tarsi brown. Fore coxa from the front with numerous white hairs and several strong yellow setae. Middle and hind coxae from the outside with several yellow cilia. Fore and hind femora with very short light and dark ventral hairs, middle femora with row of dark and light ventral cilia, at most equal in length to femora diameter. Fore tibia with 1 posteroventral seta at basal 1/3, half as long as tibia length. Fore basitarsus ventrally flattened, with dense pale pile. Length ratio of fore coxa to femora to tibia to tarsus (segments from first to fifth), 40 : 70 : 58 : 39 : 25 : 18 : 10 : 8. Middle tibia with 2 anterodorsal and 2 or 3 apical setae. Length ratio of middle coxa to femora to tibia to tarsus (segments from first to fifth), 28 : 85 : 98 : 77 : 29 : 18 : 10 : 7. Hind tibia with 1 anterodorsal, 2 short dorsal and 2 or 3 apical setae. Length ratio of hind coxa to femora to tibia to tarsus (segments from first to fifth), 20 : 100 : 140 : 64 : 36 : 24 : 12 : 7.
Wings transparent, veins brown. R1 5/12 as long as wing. R2+3 almost parallel to costa in apical half. R4+5 gently curved to M1 in apical fourth. M1+2 and M1 form the right angle. Ratio of parts of costa between R2+3 and R4+5 to those between R4+5 and M1, 16 : 3. M2 present as short stub vein and faint fold on membrane. Crossvein m-cu straight, oblique. Ratio of crossvein m-cu to apical part of M1+2 (fork-handle) to apical part of CuA1, 30 : 43 : 16. Anal vein weak, anal lobe present. Anal angle acute. Alula small but distinct. Lower calypter yellow, with brown edging and dark cilia. Halters yellow, halter stem a little longer than knob.
Abdomen metallic dark-green, with short black hairs and long black marginal setae.
Sternite with short sparse hairs. Hypopygium green-black. 8th segment positioned symmetrically, covering epandrium basodorsally and having several long setae. Cercus orange, bilobate; dorsal lobe with apical brush of long yellow undulate setae, as long as epandrium; ventral lobe twice longer than dorsal, slightly widened apicad, with black flattened setae. Basoventral lobes of both cerci forming "Organ X", fused along entire length, with longitudinal suture, long, with 2 short ventral setae.
Female. Unknown.
Length: body 3.3 mm; antenna 1.1 mm; wing-length 3.3 mm; wing-width 1.0 mm.
Distribution. Madagascar.
Etymology. The species is named for the collector, A. Pauly.
Diagnosis. B. paulyi is related to B. deconinckae, strongly differing in bilobate cercus with long thick setae, long posteroventral seta at basal third of fore tibia, one pair of long acrostichal setae. Antenna mostly black, scape mostly yellow. 8th segment positioned symmetrically, covering epandrium basodorsally. "Organ X" simple, almost bare.
9. Condylostylus basovi sp.n. (Fig. 8)
Holotype [in alcohol]. ?, Madagascar: Tam, Morarano-chrome, 25.V.1992, A. Pauly.
Paratypes [in alcohol]: 51 ??, Madagascar: 25 km W. Morarano-chrome, forét, XI.1991, bac jaune. A. Pauly. 9 ??, Madagascar: Tam, Morarano-chrome, XII.1991, A. Pauly, forét, 25 km W. ? in glycerol. Madagascar: Tam, W. Morarano-chrome, 16.V.1991, A. Pauly, forét, bac jaune.
Description. Frons metallic blue-green, shining. A strong front vertical bristle (broken) arising from small mound; postocular setae black, short. Ventral postcranium covered with irregular white hairs. Face greenish-black, silvery-white pollinose, narrow. Bulging clypeus 2/3 as long as epistome. Proboscis dark-brown; palpi black, with black hairs. Antennae black, as long as height of head. Pedicel with short dorsal and ventral bristles. First flagellomere rounded, as long as high, with short hairs. Arista dorsal, microscopically haired. Length ratio of scape to pedicel to first flagellomere to arista, 5 : 5 : 5 : 60.
Mesonotum and scutellum metallic blue-green. Pleura bronze-green, white pollinose. Five dorsocentral bristles (broken in holotype) gradually decreasing in size anteriorly with two strongest posterior pairs. Acrostichals short, biseriate. Scutellum with two pairs of strong setae (broken in holotype).
Legs mostly black-brown. Coxae black. Fore coxa at apex, trochanter, tibia and basitarsus yellow; fore femora brown in the middle, with yellow apices; middle trochanter, tibia and basitarsus dark-brown. Legs otherwise black. Fore coxa from the front with numerous yellow hairs and 3 black subapical setae. Middle coxa from the outside with light hairs and cilia. Hind coxa with one external seta. Femora without strong or long setae. All femora with light ventral hairs in basal half, half as long as femora diameter. Fore tibia without seta. Fore basitarsus 4 times longer than wide, flattened and widened except basal two fifths, with ventral pile, posteroventral row of hooked setulae in basal two fifths, row of elongate setulae along dorsal margin of flattened part and 1 or 2 short apicodorsal setae. Length ratio of fore coxa to femora to tibia to tarsus (segments from first to fifth), 50 : 63 : 75 : 53 : 22 : 13 : 9 : 7. Middle tibia with 3 short posterodorsals in basal half and 2 short ventral setae in apical half. Third and fourth tarsomeres with elongate setulae; fourth and fifth tarsomeres slightly enlarged and flattened. Length ratio of middle coxa to femora to tibia to tarsus (segments from first to fifth), 25 : 50 : 103 : 82 : 20 : 15 : 14 : 10. Last tarsomeres of hind tarsi slightly thickened. Length ratio of hind coxa to femora to tibia to tarsus (segments from first to fifth), 25 : 105 : 153 : 72 : 23 : 18 : 12 : 9.
Wings widened apically, hyaline in basal part, dark in apical half, brownish along veins, veins brown. R4+5 gently curved to M1 in apical fifth. M1+2 straight in basal half, slightly convex anteriad in middle part. M1 with nearly right-angular elbow, forming right angle with M1+2 and M2. Ratio of parts of costa between R2+3 and R4+5 to those between R4+5 and M1, 33 : 4. Crossvein m-cu straight. Ratio of crossvein m-cu to apical part of M1+2 (fork-handle) to apical part of CuA1, 26 : 49 : 15. Anal vein and lobe reduced. Anal angle absent. Lower calypter brownish, with fine cilia. Halter yellow.
Abdomen thin and long, metallic green-black, with short black hairs. Unmodified segments combined 3 times as long as mesonotum. Hypopygium black, with short black hairs. Cercus brown, long, filiform, slightly broadened at base, without basoventral lobe, with numerous black hairs; the ventral hairs at base somewhat hooked. Cercus 5 times longer than epandrium. Surstylus and epandrial lobe greatly reduced.
Female unknown.
Length: body 5.3 mm; antenna 1.1 mm; postabdomen 2.1 mm; wing-length 3.7 mm; wing-width 1.1 mm.
Distribution: Madagascar.
Etymology. The species is named for Russian dipterologist V.M. Basov.
Diagnosis. Males of C. basovi are included into the group of species with normal female-type venation of wings, differing from other species in long filiform cercus without basal swelling, simple tibiae without remarkable setae or hairs, and other characters.
10. Condylostylus burgeoni Parent (Fig 9)
Type material examined. Holotypus, ? [red label] / Musee du Congo, Kivu: Tshibinda, 18/26.XI.1932, L. Burgeon / R. Det. Z. 2966 / Condylostylus burgeoni n.sp. Type. O. Parent.
Additional material. 3 ??, Urundi, Bururi, 4.XII.1950, 2000 m [20.XII.1948, 1950 m], F.J. Francois / R.I.Sc.N.B. I.G. 24.452.
Diagnosis. C. burgeoni is closely related to C. galinae, differing in brown-black middle and hind legs and quantitative characters as follows: apical part of M1+2 (fork-handle) at least twice as long as m-cu; cercus nearly twice as long as 7th tergite and epandrium combined, with long setae from base to apex. Frons usually bare in both sexes, with strong vertical seta and at most one fine hair on small mound.
Distribution. Congo (Kinshasa), Tanzania, Kenya, Ruanda, Burundi.
11. Condylostylus chaineyi sp.n. (Fig. 10)
Holotype. ?, Fort Daufin, R.P. / Institut Scientifique Madagascar [RINS].
Paratypes [in alcohol]: 1 ?, Madagascar: Tam, Morarano-chrome, 115.VIII.1991, A. Pauly, forét, 25 km W. 3 ??, Madagascar: Morarano-chrome, 115.IX.1991, A. Pauly, forét, 25 km W. 1 ?, Madagascar: Tam, Morarano-chrome, VII.1991, A. Pauly col., forét, 25 km W. 14 ??, Madagascar: Tam, Morarano-chrome, XII.1991, A. Pauly col., forét, 25 km W. 62 ??, Madagascar: 25 km W. Morarano-chrome, forét, XI.1991, bac jaune. A. Pauly. 17 ??, Madagascar: Tam, Morarano-chrome, A. Pauly, X.1991. 1 ?, Madagascar: Tam, Morarano-chrome, 25.V.1992, A. Pauly col. 8 ??, Madagascar: Tam, W. Morarano-chrome, 16.V.1991, A. Pauly, forét, bac jaune. [RINS].
Description. Frons shining blue-violet, pollinose at base of antennae. A strong front vertical bristle bends forward, arising from small bare mound; strong postvertical bristle is positioned as a linear continuation of the postocular setal row. Ventral postcranium covered with irregular white hairs. Face black, white pollinose, narrow, 17 times as high as wide in the middle and 4 times as high as wide under antennae. Slightly bulging clypeus half as wide as epistome under antennae. Proboscis dark-brown, palpus black, with short hairs and 1 or 2 fine black setae. Antenna black, nearly as long as height of head. Pedicel with short dorsal and ventral bristles. First flagellomere rounded, as long as high, with short pubescence. Arista dorsal, microscopically haired. Length ratio of scape to pedicel to first flagellomere to arista, 4 : 4 : 5 : 55.
Mesonotum and scutellum shining green-violet, slightly pollinose. Pleura bronze-green, white pollinose. 5 dorsocentral setae gradually decreasing in size anteriorly (most setae broken in holotype). Short acrostichals in two rows. Scutellum with two pair of strong bristles, with somewhat smaller lateral setae.
Legs. Fore coxa and leg yellow, tarsus brownish; middle and hind coxae black, whitish pollinose; middle femora mostly black-brown, yellow in basal third or half; middle and hind trochanters yellow; middle tibia yellow-brown; hind femora mostly black, yellow at base; hind tibia black-brown; middle and hind tarsi black. Fore coxa from the front with yellow hairs and 3 strong black setae at apex. Middle and hind coxae from the outside with sparse yellow hairs. Hind coxa with fine yellow seta at base. Femora almost bare. Fore tibia with fine apicoventral seta. Fore basitarsus flattened and widened except basal fifth, 4 times longer than wide, with ventral pile and elongate apicodorsal setulae. Length ratio of fore coxa to femora to tibia to tarsus (segments from first to fifth), 30 : 65 : 85 : 60 : 23 : 12 : 8 : 7. Middle tibia with 1 anterodorsal, 2 or 3 short posterodorsal and 2 or 3 apical setae. 3rd and 4th tarsomeres of middle tarsus somewhat thickened, with elongate setulae. Length ratio of middle coxa to femora to tibia to tarsus (segments from first to fifth), 30 : 90 : 122 : 92 : 24 : 15 : 10 : 9. Hind tibia with inconspicuous ordinary setae, 1 or 2 short apical setae; basitarsus with 1 basoventral short seta. Last tarsomeres of hind tarsi slightly thickened. Length ratio of hind coxa to femora to tibia to tarsus (segments from first to fifth), 40 : 110 : 158 : 80 : 25 : 15 : 10 : 8.
Wings mostly hyaline, darkened anteriorly in apical half; veins brown, undisturbed. R4+5 gently curved to M1 in apical fifth. M1+2 slightly curved posterad in apical part. M1 with nearly right-angular elbow, forming right angle with M1+2 and M2. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1, 29 : 4. Crossvein m-cu straight. Ratio of crossvein m-cu to apical part of M1+2 (fork-handle) to apical part of CuA1, 25 : 41 : 13. Anal vein and lobe reduced. Anal angle absent. Lower calypter yellow, with light cilia. Halters yellow-brown, halter stem thin and long, nearly twice longer than knob.
Abdomen thin and long, mostly black-violet, with short black hairs. First two segments mostly metallic blue-green; first tergite with broad membranous excavation, longitudinal dorsal furrow and several long white lateral hairs. Unmodified segments combined 3 times longer than mesonotum. 5th and 6th segments swollen ventrally; 7th segment short. Hypopygium black, with short black hairs. Cercus black, strongly swollen at base, narrowed apicad, 6 times longer than epandrium, laterally with a row of long black setae. Surstylus and epandrial lobe greatly reduced.
Female unknown.
Length: male body 4.8 mm; antenna 0.9 mm; postabdomen 1.3 mm;
wing-length 3.5 mm; wing-width 1.05 mm.
Distribution. Madagascar.
Etymology. The species is named for English dipterologist John Chainey.
Diagnosis. Males of C. chaineyi are included into the group of species with normal female-type venation of wings, differing from other species in the following complex of characters: anterior four tibia without long setae, hind tibia without sicatrix, pedicel with comparatively short setae; cercus simple, swollen at base, strongly narrowed apicad; basoventral lobe of cercus undeveloped.
12. Condylostylus congensis Curran
Type material examined. Holotypus, ? [red label] / Musee du Congo, Mayumbe Lemba, 110.XII.1915, R. Mayne / D. Det. I 1954 / Type. Condylostylus congensis Curran.
Additional material. 1 ?, 1?, Kenya: Ongata Longai, 29.X.1995, T. Kanasugi [NMK].
Diagnosis. Similar to C. pateraeformis except as noted. Face 9 times as high as wide in the middle; legs mostly brown-black, fore coxa in apical half and fore tibia yellow; fore femora and basitarsus and sometimes middle tibia yellowish-brown; ratio of fore tibia to length of basitarsus to width of the same joint to second tarsomere, 98 : 53 : 13 : 25; fore basitarsus usually shorter than 2nd to 5th tarsomeres combined. M1+2 and M1 form the right angle; M1 nearly straight; cercus nearly twice as long as 7th tergite and epandrium combined, with egg-shaped ventral lobe.
Remark. C. congensis from Madagascar determined by P. Vanschuytbroeck belongs to C. skufjini.
Distribution: Congo (Brazzaville), Cameroons, Congo (Kinshasa), Uganda, Tanzania, Kenya, South Africa.
13. Condylostylus galinae Grichanov (Fig. 11)
Type material examined. Holotype. ?, Uganda: Ruwenzori Range, XII.1934I.1935. B.M.E.Afr. Exp. B.M.1935203 / Namwamba Valley, 6500 ft (F.W.Edwards) / Condylostylus galinae Grichanov. Paratypes. 2 ?? and 1 ?, the same labels.
Additional material. ?, Congo Belge: P.N.A., 5.II.1953, P. Vanschuytbroeck & J. Kekenbosch, 203334 / Massif Ruwenzori, Kalonge, 2010 m, Riv. Kamahoro affl. Butahu.
Diagnosis. Males of C. galinae are closely related to C. burgeoni and can be separated by the following combination of attributes: fore femora, fore and middle tibiae yellow, middle and hind femora mostly yellow; wing fork-handle 1.5 times as long as m-cu; cercus nearly thrice as long as seventh tergite and epandrium combined, with long bristles in basal half. Female is probably similar to C. burgeoni.
Distribution: West Uganda, Congo (Kinshasa).
14. Condylostylus imitator Curran
Material examined. 1 ?, Kenya: Taita hills, Chawia forest, 12.2.1997 [NMK]. 6 ??, Urundi, Rumonge, alt. 780 m, 20.II.1949 [Kitaba, 11.II.1950, 1700 m; Terr. de Rutana, 10.XI.1951, 1400 m and 25.XII.1951, 1850 m; Terr. de Kitega, 13.I.1952, 1750 m], F.J. Francois / R.I.Sc.N.B. I.G. 24.452.
Diagnosis. Similar to C. pateraeformis except as noted. Face 7 times as high as wide in the middle; legs mostly yellow, middle and hind coxae, 2nd5th tarsomeres of fore tarsus, apex of middle tibia, middle and hind tarsi, middle femora in apical fifth black-brown, hind tibia brown; ratio of fore tibia to length of basitarsus to width of the same joint to second tarsomere, 95 : 50 : 12 : 28; fore basitarsus usually shorter than 2nd to 5th tarsomeres combined. M1+2 and M1 form the right angle; M1 with distinct elbow; cercus thin, nearly thrice as long as 7th tergite and epandrium combined, ventral lobe of cercus with pointed apex.
Distribution. Congo (Kinshasa), Kenya, Tanzania, Burundi, Malawi, Zimbabwe, Mozambique, South Africa, Angola, Namibia.
15. Condylostylus kivuensis Vanschuytbroeck
Type material examined. Holotypus, ? [red label] / Coll. Mus. Congo, Kamogobe (Sud Masisi), 4.III.1936, L. Lippens (4849) / P. Vanschuytbroeck det. 195? Condylostylus kivuensis n.sp.
Diagnosis. C. kivuensis is rather similar to C. congensis and C. imitator, differing by legs entirely black-brown, with only fore trochanter and knee yellowish. M1+2 and M1 form right angle. Fore and middle tibia without very long setae. Fore basitarsus longer than 2nd to 5th tarsomeres combined. Cercus 4 times longer than epandrium. Body length 7 mm.
Distribution. Congo (Kinshasa).
16. Condylostylus paricoxa Parent
Material examined. 1 ?, Kenya: Tavara, Kitovo Forest, 24.5.1985, J. Muhangani [NMK].
Diagnosis. C. paricoxa is the only species in Africa with two long preapical setae on the male fore tibia (Grichanov, 1996c). It can be separated by the following combination of attributes: frons with white hairs; fore tibia with a long black preapical ventral hair, with 2 long thick black apicoventral setae of equal length, half as long as fore basitarsus. First tarsomere slightly swollen in middle half, with short dense ventral hairs and one thin apicodorsal seta. Cercus yellow, short, spoon-shaped, curved, especially on apex, with yellow hairs. Surstylus greatly reduced, narrow, epandrial lobe very short, with strong setae. Females differ in having a mostly yellow middle coxa and entirely yellow hind coxa.
Distribution. Kenya, Zimbabwe, Tanzania.
17. Condylostylus selitskayae sp.n. (Fig. 12)
Holotype. ?, Coll. R.I.Sc.N.B., Congo belge: Kasai, Port Francqui, 18.I.1958, F. Francois.
Description. Frons metallic blue-green, shining. A strong front vertical bristle bends forward, arising from small bare mound; shorter postvertical bristle is positioned as a linear continuation of the postocular setal row. Ocellar tubercle with a pair of strong setae and pair of hairs. Ventral postcranium covered with irregular white hairs. Face greenish-black, silvery-white pollinose, narrow, 10 times as high as wide in the middle and 4 times as high as wide under antennae. Bulging clypeus nearly half as wide as epistome under antennae. Proboscis black, palpi black, with black hairs. Antennae black, nearly as long as height of head. Pedicel with short dorsal and ventral bristles. First flagellomere rounded, as long as high, with short hairs. Arista dorsal, microscopically haired. Length ratio of scape to pedicel to first flagellomere to arista, 5 : 5 : 5 : 60.
Mesonotum and scutellum metallic green, slightly pollinose. Pleura bronze-green, white pollinose. Five dorsocentral bristles gradually decreasing in size anteriorly with two strongest posterior pairs. Short acrostichals in two rows, restricted to anterior 2/3 of mesonotum. Scutellum with two pairs of strong setae of approximately equal length.
Legs mostly black. Fore coxa mostly black, yellow in apical 1/3; fore trochanter yellow; fore femora brown in basal half, yellow in apical half; fore tibia mostly yellow, progressively brown in apical half; fore basitarsus brown; middle coxa at apex, trochanter and knee brownish. Fore coxa from the front with numerous yellow hairs and 3 black subapical setae. Middle coxa from the outside with light hairs and black cilia. Hind coxa with one black external seta. Femora without strong or long setae. Fore and hind femora with posteroventral hairs, half as long as femora diameter. Fore tibia with short black subapical ventral seta. Fore basitarsus 4 times longer than wide, flattened and widened except basal fifth, with ventral pile and 2 short apicodorsal setae. Length ratio of fore coxa to femora to tibia to tarsus (segments from first to fifth), 50 : 85 : 90 : 47 : 22 : 14 : 9 : 10. Middle tibia with 2 short posterodorsals in basal half and several apical setae, and erect ventral setulae in apical half. Third and fourth tarsomeres with elongate setulae; fourth and fifth tarsomeres slightly enlarged and flattened. Length ratio of middle coxa to femora to tibia to tarsus (segments from first to fifth), 40 : 100 : 125 : 70 : 21 : 15 : 13 : 10. Last tarsomeres of hind tarsi slightly thickened. Length ratio of hind coxa to femora to tibia to tarsus (segments from first to fifth), 25 : 130 : 175 : 75 : 25 : 17 : 11 : 10.
Wings widened apically, mostly hyaline, with brownish stripe along costa in apical half and smoky spot at m-cu, veins brown. R4+5 gently curved to M1 in apical fifth. M1+2 convex posteriad in basal half, convex anteriad in apical half. M1 slightly convex basad, forming right angle with M1+2. M2 short, forming wide arc with M1. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1, 75 : 8. Crossvein m-cu straight. Ratio of crossvein m-cu to apical part of M1+2 (fork-handle) to apical part of CuA, 21 : 83 : 29. Anal vein and lobe reduced. Anal angle absent. Lower calypter brownish, with fine, shining light, cilia. Halter brown, halter stem thin, 1.5 times longer than knob, with row of setulae in front of knob.
Abdomen thin and long, metallic green-black, posteriorly entirely black, with short black hairs. First tergite with broad membranous excavation, longitudinal dorsal furrow and short white lateral hairs. Unmodified segments combined 3 times as long as mesonotum. Hypopygium black, with short black hairs. Cercus long, filiform, swollen at base, black, with distinct hairy inner apophysis at base and numerous black hairs along entire length. Cercus nearly 6 times as long as epandrium. Surstylus and epandrial lobe greatly reduced.
Female unknown.
Length: body 5.3 mm; antenna 1.15 mm; postabdomen 1.8 mm; wing-length 4.1 mm; wing-width 1.4 mm.
Distribution: Congo (Kinshasa).
Etymology. The species is named for Russian entomologist Oksana Selitskaya.
Diagnosis. Males of C. selitskayae is related to C. congensis, differing in erect ventral setulae in apical half of middle tibia and weakly developed basoventral lobe of cercus. Fore coxa yellow in apical third; fore femora and tibia half yellow; fore basitarsus shorter than 2nd to 5th tarsomeres combined. M1+2 and M1 forming right angle. Venation abnormal: M1+2 curved towards posterior wing margin, M2 short, forming wide arc with M1.
18. Condylostylus skufjini sp.n. (Fig. 13)
Holotype. ? on pin, Fenerive, Madagascar, Dec. 1955, B. Stuckenberg / P. Vanschuytbroeck det. 1957, Condylostylus congoensis Curran / R.I.Sc.N.B. I.G. 20938 [RINS].
Paratypes [mostly in alcohol]. ? on pin, La Mandraka, XII.1951, N.S.H. Krauss / Institut Scientifique Madagascar. 17 ??, Madagascar: Tam, Morarano-chrome, A. Pauly, X.1991. 1 ?, Madagascar: Tam, Morarano-chrome, XII.1991, A. Pauly. 12 ??, Madagascar: Tam, W. Morarano-chrome, 16.V.1991, A. Pauly, forét, bac jaune. 1 ?, Madagascar: Tam, Morarano-chrome, 1630.IX.1991, A. Pauly, forét, 25 km W. 5 ??, Madagascar: Foulpointe, 11.XI.1993, A. Pauly col., bac jaune, forét. 1 ?, Madagascar: Tam, Foulpointe, X.1993, A. Pauly col. 1 ?, Madagascar: Foulpointe, X.1993, A. Pauly col., forét . 1 ?, Madagascar: Foulpointe, 2.XI.1991, A. Pauly., plage, bac jaune.[RINS].
Description. Frons metallic bluish-green, white pollinose at base of antennae. A strong front vertical bristle bends forward, arising from small bare mound; strong postvertical bristle is positioned as a linear continuation of the postocular setal row. Ventral postcranium covered with irregular white hairs. Face greenish-black, silvery-white pollinose, narrow, 12 times as high as wide in the middle and 4 times as high as wide under antennae. Bulging clypeus nearly half as wide as epistome under antennae. Proboscis black, palpus black, with light hairs and 2 fine black setae. Antenna black, as long as height of head. Pedicel with short dorsal and ventral bristles. First flagellomere rounded, as long as high, with short pubescence. Arista dorsal, microscopically haired. Length ratio of scape to pedicel to first flagellomere to arista, 4 : 4 : 5 : 67.
Mesonotum and scutellum brilliantly shining green-blue. Pleura bronze-black, white pollinose. 5 dorsocentral setae gradually decreasing in size anteriorly with two strongest posterior pairs. Short acrostichals in two rows, restricted to anterior half of mesonotum. Scutellum with two pairs of strong bristles, with lateral setae somewhat smaller.
Legs mostly yellow. Middle and hind coxae dark-brown; middle femora in apical fourth, middle tibia and 2nd to 5th tarsomeres of fore tarsus brownish; hind femora in apical third, hind tibia, middle and hind tarsi black-brown. Fore coxa from the front with yellow hairs and 2 or 3 black setae at apex. Middle and hind coxae from the outside with sparse yellow hairs. Hind coxa with fine light seta at base. Femora without strong or long bristles and hairs. Fore tibia with 1 black apicoventral seta. Fore basitarsus flattened and widened except basal third, 4.6 times longer than wide, with ventral pile and elongate apicodorsal setulae. Length ratio of fore coxa to femora to tibia to tarsus (segments from first to fifth), 46 : 66 : 80 : 55 : 19 : 11 : 6 : 6. Middle tibia with ventral row of 5 setae in apical fifth, twice longer than tibia diameter, and preceding row of several elongate setulae, otherwise bare. 3rd to 5th tarsomeres of middle tarsus slightly flattened. Length ratio of middle coxa to femora to tibia to tarsus (segments from first to fifth), 33 : 90 : 130 : 87 : 26 : 23 : 10 : 10. 1st to 4th tarsomeres of hind tarsi flattened and slightly widened. Length ratio of hind coxa to femora to tibia to tarsus (segments from first to fifth), 25 : 120 : 173 : 86 : 27 : 18 : 10 : 9.
Wings mostly hyaline, slightly darkened anteriorly at apex including base of M1; veins brown, undisturbed. R4+5 nearly straight, gently curved to M1 in apical fifth. M1+2 slightly curved posterad in apical part. M1 with nearly right-angular elbow, forming right angle with M1+2 and M2. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1, 28 : 4. Crossvein m-cu straight. Ratio of crossvein m-cu to apical part of M1+2 (fork-handle) to apical part of CuA1, 23 : 43 : 14. Anal vein and lobe reduced. Anal angle absent. Lower calypter yellow, with light cilia. Halters yellow, halter stem thin and long, twice longer than knob, with several dark setulae at the middle.
Abdomen thin and long, mostly black, with short black hairs. First two segments mostly metallic green; first tergite with broad membranous excavation, longitudinal dorsal furrow and several long white lateral hairs. Unmodified segments combined nearly 3 times longer than mesonotum. Hypopygium black, with short black hairs. Epandrium rounded. Cercus black-brown, short, widened apicad and truncated at apex, with rounded angles and short black hairs along entire surface. Cercus as long as epandrium. Surstylus and epandrial lobe greatly reduced.
Female unknown.
Length: body 4.7 5.3 mm; antenna 1.1 mm; postabdomen 0.6 mm; wing-length 3.6 mm; wing-width 1.0 mm.
Distribution. Madagascar.
Etymology. The species is named for Russian dipterologist Prof. K.V. Skufjin.
Diagnosis. C. skufjini is related to C. chaineyi, differing in short cercus and long ventral setae on middle tibia (see diagnosis of C. chaineyi).
19. Ethiosciapus bicalcaratus (Parent)
Type material examined. Holotypus, ? [red label] / Musee du Congo, Ituri: Mont Wago, 24.XI.1928, A. Collart / R. Det. R. 2413 / Sciopus bicalcaratus n.sp. Type. O. Parent.
Additional type material. Holotype ?, Congo Belge: Eala, 27.IV.1936, J. Ghesquiere / R. Mus. Hist. Nat. Belg. I.G. 10.482 / Sciopus cilifrons [nomen nudum] n.sp. Type. O. Parent / O. Parent, 1936. Sciopus cilifrons n.sp. / Type [red label] / S. setifrons sp. of Bull. Mus. Hist. Nat. Belg., XII18 (1937) p. 14, pl. 4, fig. 33, 34 / Ethiosciapus bicalcaratus (Par.), det. Grichanov.
Additional material. 1 ?, Urundi, Bururi, alt. 1950 m., VI.1948, F.J. Francois / R.I.Sc.N.B. I.G. 24.452. [RINS].
Diagnosis. Male with a group of hairs laterally on frons, 2 strong posterior and hairlike anterior dorsocentral setae; 3 long acrostichals. Fore coxa yellow except base, other coxae black; femora yellow, with long black ventral hairs, fore tibia with 2 long posteroventral bristles; fore basitarsus ventrally flattened. Lower calypter with dark, though shining light, cilia. Third sternite with 2 strong black setae. Cercus yellow, with brown margin on apex, pale ventral hairs and black lateral and apical hairs, with thin basoventral hook.
Remark. Holotype of Sciapus bicalcaratus is identical to holotype of Sciapus setifrons.
Distribution. Congo (Kinshasa), Uganda, Burundi, Madagascar, Comores, St. Helena.
20. Ethiosciapus exarmatus (Parent)
Type material examined. Holotypus, ? [red label] / Musee du Congo, Ituri: Missa Moke (Faradji), 20.II.1930, A. Collart / R. Det. O. 2413 / Sciopus exarmatus n.sp. Type. O. Parent.
Diagnosis. Males with group of hairs laterally on frons; femora with long black ventral hairs; 3 long acrostichal setae; alula well developed. All coxae black. Femora mostly yellow. Fore femora black ventrally in basal half, hind femora black on apex. Fore tibia with at most 2 long posteroventrals. Cercus with short setae, without distinct groups of hairs, with strong pointed and strongly curved basoventral hook.
Remark. E. exarmatus determined by P. Vanschuytbroeck (male and female
from Madagascar are examined, RINS) belongs to Amblypsilopus cilifrons, although I have seen several males from Madagascar (RINS), quite similar to this species.
Distribution. Congo (Kinshasa), ?Madagascar.
21. Ethiosciapus finitimus (Parent) (Fig. 14)
Type material examined. Holotype, ?, Congo Belg.: Rutshuru, XII.1936, J. Ghesquiere, 3566 / R. Mus. Hist. Nat. Belg. 10.482 / O. Parent det., 1937: Sciopus finitimus [nomen nudum]. Type. O. Parent.
Additional type material. Holotype, ?, Uganda: Ruwenzori Range, XII.1934I.1935. B.M. E. Afr. Exp. B.M. 1935203 / Kilembe, 4500 ft., F.W. Edwards / Holotype Ethiosciapus skufjini Grichanov; paratype, ?with the same labels.
Additional material. 1 ?, Congo Belge: P.N.A., 2628.VIII.1953, P. Vanschuytbroeck & V. Hendricks 49995005 / Secteur Tshiaberimu, Riv. Mbulikerere, affl. dr. Talia N, 2720 m [RINS].
Diagnosis. E. finitimus is close to E. latipes, differing in longer ventral setae on all femora, black halters, other colour characters, and hypopygium morphology.
Remark. E. finitimus was described as Chrysosoma (Sciopus?) finitimus Parent (1939). So, the name Sciopus finitimus in this paper is nomen nudum. Holotype of C. finitimus is identical to holotype of E. skufjini.
Distribution. Congo (Kinshasa), Uganda.
22. Ethiosciapus flavirostris (Loew)
Material examined. 1 ? [in glycerol], Madagascar: Fia Ambalamanakana, 13.III.1994, A. Pauly [RINS].
Diagnosis. Male with a group of hairs laterally on frons, 2 strong posterior and hairlike anterior dorsocentral setae; 3 long acrostichals. Fore coxa yellow except base, other coxae black; femora yellow, with long black ventral hairs, fore tibia with 2 long posteroventral bristles; fore basitarsomere ventrally flattened. Lower calypter with light cilia. Third sternite with 2 strong black setae. Cercus yellow, with brown margin on apex, pale ventral hairs and black lateral and apical hairs, with thin basoventral hook. E. integer is possible synonym to this species. E. flavirostris is very closely related to E. bicalcaratus.
Distribution. South Africa, Mozambique, Madagascar, ?Ethiopia.
23. Ethiosciapus inflexus (Becker)
Material examined. 1 ?, Kenya: Kakamega bridge, Isiukhu river, 17.IX.1995, N 0º16'46", E 34º46'61", Leg. Earthwatch Team 5 [NMK]. 3 ??, Urundi, Bururi, alt. 1950 m, III.1948 [Terr. de Kitega, 23.II.1952, 1750 m], F.J. Francois / R.I.Sc.N.B. I.G. 24.452. 1 ?, Costermansville, Congo Belge, J. Wolfs / R.I.Sc.N.B. 24.236, Coll. M. Bequaert. 11 ??, Madagascar: Tam, Morarano-chrome, X.1991, A. Pauly. 20 ??, Madagascar: 25 km W. Morarano-chrome, forét, XI.1991, bac jaune. A. Pauly. 3 ??, Madagascar: Tam, Morarano-chrome, XII.1991, A. Pauly, forét, 25 km W. 12 ??, Madagascar: Tam, Morarano-chrome, 1630.IX.1991, A. Pauly, forét, 25 km W. 1 ??, Madagascar: Tam, Manakambahiny Atn., 1825.I.1991, A. Pauly col., forét. 3 ??, Madagascar: Tam, Manakambahiny Atn., 1723.III.1991, A. Pauly, forét. 3 ??, Madagascar: Tam, Morarano-chrome, 25.V.1992, A. Pauly. 28 ??, Madagascar: Tam, Morarano-chrome, VII, 115.VIII and 115.IX.1991, A. Pauly., forét, 25 km W. [RINS].
Diagnosis. Male with a group of hairs laterally on frons, 2 strong posterior and hairlike anterior dorsocentral setae; 3 long acrostichals. All coxae black; femora mostly black, with long black ventral hairs, fore tibia with 1 to 3 long posteroventral bristles; fore basitarsus ventrally flattened. Lower calypter with black cilia. Third sternite with long black bristles; first tergite and sternite with long white hairs. Cercus yellow, strap-like, slightly broadened towards apex, with dense pale ventral hairs, a row of black lateral bristles, and sclerotized pointed basoventral hook.
Remark. Descriptions of E. inflexus and E. dilectus have no significant difference and the two species are possible synonyms. Large series of males collected on Madagascar in yellow water traps displays a great variability of specimens in anterior four femora coloration (black in basal 1/3 to 4/5) and number of long posteroventral setae on fore tibia (1 to 3).
Distribution. Tanzania, Uganda, Kenya, Burundi, Congo (Kinshasa), South Africa, Madagascar, St. Helena..
24. Ethiosciapus latipes (Parent)
Material examined (all in alcohol). 2 ??, Madagascar: Tam, Morarano-chrome, XII.1991, A. Pauly, forét, 25 km W. 10 ??, 1 ?, Madagascar: Morarano-chrome, VII, 115.VIII, 115, 1630.IX.1991, A. Pauly., forét, 25 km W. 2 ??, Madagascar: Tam, Morarano-chrome, 25.V.1991, A. Pauly. 26 ??, Madagascar: Tam, Morarano-chrome, X.1991, A. Pauly. About 300 ??, Madagascar: 25 km W. Morarano-chrome, forét, XI.1991, bac jaune. A. Pauly. 4 ??, Madagascar: Tam, W. Morarano-chrome, forét, 16.V.1991, bac jaune. A. Pauly. 1 ?, Madagascar: Tam, Manakambahiny Atn., 1723.III.1991, A. Pauly, forét. 2 ??, Madagascar: Foulpointe, 11.XI.1993, A. Pauly col., bac jaune, forét.
Diagnosis. E. latipes is closely related to E. finitimus, differing in shorter setae on femora (a little longer than diameter of fore and hind femora, half as long as midfemora diameter). Despite the description of this species by Parent, frons has a group of lateral hairs; 4th and 5th segments of hind tarsus slightly broadened and flattened; halters brown. All coxae black; hind femora brown at apex; fore tibia with 2 or 3 moderately long posteroventral setae. 3rd abdominal sternite without strong setae. Cercus with short lateral setae; these setae stronger, than corresponding setae in E. finitimus, and cercal basoventral hook somewhat smaller.
Distribution. Madagascar.
25. Bickeliolus alluaudi (Parent)
Material examined [all in alcohol]. 14 ??, Madagascar: Tam, Morarano-chrome, XII.1991, A. Pauly col., forét, 25 km W. 1 ?, Madagascar: Tam, Manahambahiny Atn, 1825.I.1991, A. Pauly col., forét. 1 ?, Madagascar: Tam, Morarano-chrome, VII.1991, A. Pauly col., forét. 1 ?, Madagascar: Tam, Morarano-chrome, 25.V.1992, A. Pauly col. About 80 ??, Madagascar: 25 km W. Morarano-chrome, forét, XI.1991, bac jaune. A. Pauly. 14 ??, Madagascar: Ambatondvaraha, 26.II.1991, A. Pauly col. [RINS].
Diagnosis. B. alluaudi is very close to B. trochanteralis from South Africa, differing in suboval, rather than subtriangular, basoventral process of cercus. Male with strong vertical seta; acrostichals very short. Antenna black. Legs including trochanters yellow, middle trochanter with a fringe of long yellow ventral hairs, middle femora with a few fine yellow ventral hairs at base, fore basitarsus ventrally flattened. Cercus narrowed in the middle, with apical brush of long setae and suboval basoventral sclerotized process on thin stem.
Distribution. Madagascar.
26. Bickeliolus haemorhoidalis (Becker)
Material examined. 2 ??, Urundi: Rumonge, alt. 780 m, 15.X.1948 [26.XI.1948], F.J. Francois / R.I.Sc.N.B. I.G. 24.452; 1 ?, Urundi: Terr. de Bubanza, 23.VI.1953, F.J. Francois / Colline: Kagunuzi (Imbo), alt. 900 m. [RINS].
Diagnosis. Male with strong vertical seta, 4 strong dorsocentrals and microscopic acrostichals. Antenna black. Legs yellow; middle and hind coxae black; fore coxa yellow, with 4 long thick yellow setae in middle and 1 yellow apical seta of glued hairs. All femora with white ventral hairs at base, longer than femora diameter; fore basitarsus ventrally flattened, with pale pile. Cercus practically bilobate; ventral lobe short, wide, with ventral spine-like process; dorsal lobe thin, twice longer than ventral lobe, slightly widened at apex, with subapical ventral brush of short hairs and apical brush of long hairs. Basoventral sclerotized process club-shaped, truncate at apex, with strong apical setae.
Distribution. Ethiopia, Uganda, Burundi, South Africa.
27. Bickeliolus lamellatus (Parent)
Type material examined. Holotypus, ? [red label] / Musee du Congo,
Rutshuru, I1934, Dr. De Wulf / R. Det. CC.2966 / Sciopus lamellatus n.sp. Type. O. Parent.
Additional material. 1 ?, N'Goma (Kivu), 17/19.IX.1935, Dr. H. Damas, Parc Nat. Albert / P. Vanschuytbroeck det. 1950, Sciopus rectangularis Parent; 7??, 1?, Congo Belge: P.N.A., 2628.VIII.1953 [26.X.1953; 29.VIII.1956; 10.VIII.1957; 6.IX.1956; 18.XI.1956], P. Vanschuytbroeck & V. Hendricks / Secteur Tshiaberimu, Riv. Mbulikerere, affl. dr. Talia N, 2720 m [Massif Ruwenzori, Kiurama, 2100 m; Secteur Nord, riv. Mukandwe affl. dr. Talia, 1200 m; Secteur Nord, Kamusonge, massif pres Mutsora, 1420 m; Massif Ruwenzori, riv. Lume (moyenne) affl. Semliki, 1830 m]; 2??, 5??, Congo belge: Kivu [P.N.A.], Rutshuru [Nyongera-Butumba; Lubirizi; Riv. Fuku], 20 au 21.XII.1933 [6 au 8, 18 au 23.VI.1934; 517.VII.1935], 12181285 m, G.F. de Witte: 1669 [131, 431, 448, 1622, 1645, 1662]. [RINS].
Diagnosis. B. lamellatus is closely related to B. maslovae. Male with strong vertical seta and 4 strong dorsocentral bristles, the anteriormost short; acrostichals very short. Antenna black. Legs including trochanters yellow, fore and middle femora with short white ventral hairs, fore basitarsus ventrally flattened. Cercus tapering, with apical brush of long hairs and subtriangular basoventral sclerotized process.
Distribution. Congo (Kinshasa), Ruanda, Uganda, Tanzania, St. Helena.
28. Bickeliolus lutescens (Vanschuytbroeck)
Type material examined. Paratype ?, Ambila, VII.51, R.P. Forét cotiere / Institut Scientifique Madagascar / R.I.Sc.Nat.Belg. I.G. 18441 / Paratype / P. Vanschuytbroeck det. 195? Chrysosoma lutescens n.sp. [RINS].
Additional material (all in alcohol). About 95 ??, Madagascar: Foulpointe, XI.1995, A. Pauly rec., plage, bac j. About 100 ??, Madagascar: Foulpointe, 2.XI.1991, A. Pauly., plage, bac j. 7 ??, Madagascar: Foulpointe, 11.XI.1993, A. Pauly col., bac jaune, forét. 1 ?, 4 ?, Madagascar: Foulpointe, X.1993, A. Pauly col., forét, bac j. 9 ??, Madagascar: Foulpointe, XI.1995, A. Pauly. [RINS].
Diagnosis. Scape yellow, other articles dark brown dorsally; first flagellomere triangular, with rounded apex. Anterior coxa yellow, black at extreme base, with yellow setae; middle and hind coxae black. Legs yellow except apical tarsomeres. Femora bare. Fore tibia with 1 dorsal seta at base, 1 long posterior seta at basal 1/3. Middle tibia with 1 dorsal, 2 anterior setae. Hind tibia with 1 anterodorsal, 2 posterodorsal, 1 anterior setae. Fore basitarsus enlarged and ventrally flattened, with pile of hairs; 5th tarsomere of the same tarsus black, slightly flattened. Hind basitarsus with 1 short basoventral seta. Cross-vein m-cu nearly straight, slightly sinuous, approximately equal to fork-handle M; CuA1 approximately 2/3 as long as m-cu. Alula developed. Anal angle sharp. Cercus approximately 5 times longer than wide at base; cercus at base twice wider than in middle; cercal basoventral hook thick, 1/5 as long as cercus; apex of cercus slightly enlarged and rounded, with spade-like fringe of yellow cilia at basal 1/3; cercal cilia entirely yellow, 1/3 or 1/4 as long as cercus.
Distribution. Madagascar.
29. Chrysosoma (Chrysosoma) aequatoriale Parent
Type material examined. Holotype, ?, Wendji, 30.VI.27, A. Collart / Prov. Equateur (Congo Belge / Chrysosoma aequatoriale n.sp. Type. O. Parent rev., 1933.
Diagnosis. Antenna red-brown; mesonotum without distinct stripes. Lower calypter with black cilia. Male middle basitarsus without long setae. All coxae black, sometimes fore coxa yellow at apex; fore femora black in basal quarter, middle femora black in basal 3/5; middle femora with black ventral setae in the middle; hind tibia black; middle tarsus whitish; 2nd to 4th tarsomeres of middle tarsus with posterodorsal fringe of white flat setae, as long as tarsomeres diameter. Cercus simple, tapering, without tooth.
Distribution. Congo (Kinshasa), Uganda.
Chrysosoma (Chrysosoma) aequilobatum Parent
Type material examined. Holotype, ?, Musee du Congo, Mayumbe Lemba, XII.1915, R. Mayne / Chrysosoma aequilobatum n.sp. Type. O. Parent.
Diagnosis. C. aequilobatum is included into the group of species with very long setae on middle basitarsus. Hind femora entirely black. Middle tibia without long setae. Middle basitarsus with two long setae, without white preapical ring, Cercus shallow bifurcated, with equal lobes.
Distribution. Congo (Kinshasa), Congo (Brazzaville).
30. Chrysosoma (Chrysosoma) albocrinitatum Curran
Type material examined. Holotypus, ? [red label] / Musee du Congo, Mayumbe: Kiniati, 7.IV.1911, R. Mayne / R. Det. G. 1155 / Holotype Chrysosoma albocrinitatum Curran.
Diagnosis. C. albocrinitatum is related to a group of species having a row of cilia on middle basitarsus; these cilia 3 or 4 times as long as tarsomere diameter. It differs from C. crinipes in second tarsomere of fore tarsus 2/3 to 3/4 as long as basitarsus, the latter swollen, with ventral pile of short hairs in basal 1/2 or 2/3 and pectination of long hooked hairs in apical half; second tarsomere also with hooked hairs along ventral side, the hairs longer than tarsomere diameter.
Distribution. Congo (Kinshasa), Congo (Brazzaville).
31. Chrysosoma (Chrysosoma) alboguttatum Parent
Material examined. 2 ??, 2 ??, Urundi: Bururi, alt. 1950 [1900] m, III.1948 [8.I.1949; 5.III.1953], F.J. Francois / R.I.Sc.N.B. I.G. 24.452[RINS]. 1 ?, LabéGuinée, 21IX981, C. Bakary.
Diagnosis. Male tibiae and tarsi without long setae, with very short erect setulae; fore coxa yellow; antenna black; wing brown with two hyaline windows and transparent posterior edge; cercus simple, short, narrow, slightly widened at apex.
Distribution. Cameroons, Guinea, Burundi.
32. Chrysosoma (Chrysosoma) arduus (Parent)
Type material examined. Holotype, ?, Congo Belge: Eala, XI.1934, J. Ghesquiere / Reg. Mus. Hist. Nat. Belg. I.G. 10.482 / Sciopus arduus n.sp. Type. O. Parent det., 1935 / Type [red label]; paratypes, 4 ??, same locality, with additional red label "cotype".
Diagnosis. Face widely bulging, clypeus slightly separated from eyes. Wings strongly maculated; m-cu strongly sinuate. Middle femora with ventral seta at base. Fore femora yellow in apical two thirds, with 1 or 2 fine black ventral setae; fore tibia with 1 anterodorsal and 2 posterodorsal, middle tibia with 3 anterodorsal, 3 posterodorsal and 2 ventral, middle basitarsus with 5 ventral setulae; hind tibia with 1 anterodorsal and 1 posterodorsal at base, 4 dorsal, 3 anterior and 34 short ventral setae; hind basitarsus with 1 basoventral seta. Despite the description by Parent (1936), the fore femora has 0 to 2 dark and 5 to 10 light fine ventral cilia approximately as long as femora diameter; these cilia arranged in two irregular rows in basal half of femora. The species is possible synonym to C. norma.
Distribution. Congo (Kinshasa).
33. Chrysosoma (Chrysosoma) bacchi Dyte
Material examined. 1 ?, Urundi: Rutana, 24.V.1950, 1800 m, F.J. Francois / R.I.Sc.N.B. I.G. 24.452 [RINS].
Diagnosis. Male middle basitarsus with white preapical ring, usually covered with very short yellow pectination on dorsal side; middle basitarsus with only three long setae; middle tibia with two long setae; hind femora black except apical quarter and almost bare.
Distribution. Tanzania, Burundi.
34. Chrysosoma (Chrysosoma) bredoi Parent
Type material examined. Holotype, ?, Musee du Congo, Lukolela, III.1929, H.J. Bredo / Chrysosoma bredoi n.sp. Type. O. Parent; 3 ??, 1 ?, Congo Belge: Eala, 7.V, IX, XI.1935, IX.1936, J. Ghesquiere / R. Mus. Hist. Nat. Belg. I.G. 10.482 / Chrysosoma bredoi Par. O. Parent det., 1936.
Diagnosis. Closely related to C. hirsutulum, differing in at most 1 or 2 long setae at base of middle tibia; middle basitarsus with 3 to 5 long setae; middle tarsus with erect pectination from the middle of 1st joint; femora dark-brown except apices. Holotype with head missing.
Distribution. Congo (Kinshasa).
35. Chrysosoma (Kalocheta) collarti (Parent)
Type material examined. Holotypus, ? [red label] / Musee du Congo, Ituri: Gaduma Mala (Faradji), 13.III.1930, A. Collart / R. Det. T.2414 / Kalocheta collarti n.sp. Type. O. Parent.
Additional material. 2 ??, 1 ?, Kenya: Kakamega forest, Ikuywa river, 25.VI.1995, N 0º12'73", E 34º55'42", Leg. Earthwatch Team 4; 2 males, 2 ??, Kenya: Kakamega forest, Buyaungu Res., 21.I, 2.XII 1995, 21.XI.1994, far end path (Salazar circ), N 0º20'84", E 34º51'93", Leg. Earthwatch Team 1, 2, 6 [NMK]; 1 ?, Kenya: Kakamega forest, nr. road betw. Rondo estate & Isecheno, 23.I.1995, Leg. Earthwatch Team 2 [NMK]; 1 ?, Kenya: Kakamega forest, path betw. Isecheno stat. & Kalunya glade, 20.I.1995, N 0º14'48", E 34º51'88", Leg. Earthwatch Team 2 [NMK].
Diagnosis. Fore coxae with white hairs and black bristles; apical and median brown spots have crosspiece near junction of M 3+4 and posterior transverse (m-cu) wing veins. Males with hairlike part of arista twice as long as strap-like part; lower calypter with black hairs; Cercus with short dorsal tooth at basal third and short thin subapical dorsal tooth. Metallic epistome and clypeus appearing white pollinose under certain lightening. See also description of female by Grichanov (1995).
Distribution. Congo (Kinshasa), Tanzania, Kenya, Uganda.
36. Chrysosoma (Chrysosoma) consentium Curran
Type material examined. Holotypus, ? [red label] / Musee du Congo, Mayumbe: Kiniati, 7.VI.1911, R. Mayne / R. Det. H. 1155 / Holotype Chrysosoma consentium Curran. [RMCA].
Additional type material. Holotype, ?, Yumbi, 1.VII.1912, Dr. P. Mouchet / Chrysosoma fortunatum n.sp. Type. O. Parent det., 1933 / Type [red label] [RINS].
Diagnosis. C. consentium is similar to C. mesotrichum and C. schoutedeni, differing mainly in morphology of hypopygium. Fore and middle femora black in basal third; hind leg entirely black; middle tibia and tarsus except apex yellow. All femora with white ventral hairs in basal half. First and second tarsomeres of middle tarsus with additional row of short dorsal hairs, which more than twice as long as tarsomere diameter; middle tibia with 7 to 10 long posterodorsal setae; middle basitarsus with 6 to 10 long setae. 2nd to 5th tarsomeres of middle tarsus with black hairs; 5th tarsomere with pile of white hairs. Cercus with short dorsal tooth, at most half as long as width of cercus at apex, with short hairs in apical half of cercus.
Remark. Despite Curran (1925), C. consentium has nothing to do with C. albocrinitatum, but is identical to the holotype of C. fortunatum.
Distribution. Congo (Kinshasa), Congo (Brazzaville).
37. Chrysosoma (Chrysosoma) continuum Curran
Type material examined. Holotypus, ? [red label] / Musee du Congo, Bas-Uele: Koteli, I.1925, Dr. H. Schouteden / Type Chrysosoma continuum Curran.
Diagnosis. C. continuum is associated with a group of species having long setae on middle tibia and basitarsus. Middle tibia with 78 long setae; middle basitarsus without white preapical ring, with 7 long setae in apical half; second to fifth tarsomeres with a row of white hairs, without black hairs. Fore femora widely black at base; hind femora mostly blackish. Antenna red. Wing vein m-cu, measured along sinuation, nearly thrice as long as fork-handle. Cercus simple, with short dorsal apophysis.
Distribution. Congo (Kinshasa), Congo (Brazzaville).
38. Chrysosoma (Chrysosoma) corruptor Parent
Type material examined. Holotypus, ? [red label] / Musee du Congo, Stanleville, 19.II.1928, A. Collart / R. Det. J. 2413 / Chrysosoma corruptor n.sp. Type. O. Parent.
Diagnosis. Holotype of C. corruptor is similar to description of. C minusculum Becker (1923). Antenna black. Mesonotum metallic, pleura and coxae with silvery tinge. Two dorsocentrals. Vein m-cu sinuate. First and second abdominal segment with white tinge (see from behind). Only fore coxa yellow. Middle and hind coxae with pale ciliation. Middle tibia and tarsus without erect pubescence. Middle basitarsus without long setae. Cercus bifurcated, with lobes more than half as long as cercus.
Distribution. Congo (Kinshasa).
39. Chrysosoma (Chrysosoma) hirsutulum Parent
Type material examined. Holotypus, ? [red label] / Musee du Congo, Equateur, Boende, 11.III.1926, R.P. Hulstaert / Chrysosoma hirsutulum n.sp. Type. O. Parent.
Diagnosis. C. hirsutulum is associated with a group of species having long setae on middle tibia and basitarsus. Middle tibia with 3 long setae (basal one shorter) in basal half; middle basitarsus without white preapical ring, with three long setae in apical half; middle tibia and tarsus with erect pectination (holotype with broken 3rd5th tarsomeres of middle tarsus). Cercus simple, with short dorsal apophysis.
Distribution. Congo (Kinshasa).
40. Chrysosoma (Chrysosoma) ituriense Parent
Type material examined. Holotypus, ? [red label] / Musee du Congo, Ituri: Genge (Nizi), 16.III.1929, A. Collart / R. Det. A.2413 / Chrysosoma ituriense n.sp. Type. O. Parent.
Diagnosis. Antenna black. Lower calypter with pale cilia. All coxae black, femora yellow. Middle tibia and basitarsus without long setae, middle tarsus without white hairs; middle tibia with erect pectination on all sides, hairs half as long as tibia diameter; 1st to 4th tarsomeres with mostly dorsal pectination. Cercus not bifurcated, short, rounded in apical half, with short hairs.
Distribution. Congo (Kinshasa), Malawi.
41. Chrysosoma (Chrysosoma) katangense Curran
Type material examined. Holotypus, ? [red label] / Musee du Congo, Lubumbashi, Mt. Katanga, 1.XI.21, Dr. M. Bequaert / R. Det. D. 1155 / Holotype Chrysosoma katangensis Curran.
Additional type material. Holotypus, ? [red label] / Musee du Congo, Elisabethville, 1927, Dr. M. Bequaert / R. Det. H. 2412 / Chrysosoma speciosum n.sp. Type. O. Parent.
Diagnosis. C. katangense is associated with a group of species having black antenna and numerous long setae on middle tibia and basitarsus. It differs from other species in sinuous vein m-cu, middle basitarsus with four or five long setae and row of shorter setae, half as long as the longest setae; middle tibia with 1 very long apical seta and 3 or 4 long setae, half as long as the longest setae on basitarsus; middle basitarsus dark or yellow, without white preapical ring covered with yellow pectination; second to fifth tarsomeres of middle tarsus without remarkable ciliation; cercus not bifurcated, rounded on apex, with dorsoapical excavation and short pointed apophysis at the middle.
Remark. Holotype of C. speciosum Parent has no difference from holotype of C. katangense Curran.
Distribution. Congo (Kinshasa), Sudan.
42. Chrysosoma (Chrysosoma) lavinia Curran
Type material examined. Holotypus, ? [red label] / Musee du Congo, Bas-Uele: Koteli, 121.I.1925, Dr. Schouteden / Type Chrysosoma lavinia Curran.
Additional material. 6 ??, Urundi, For. M'Bururi, alt. 2000 m, 4.IX.1948 [3.IX.1949; 10.X.1948], F.J. Francois / R.I.Sc.N.B. I.G. 24.452.
Diagnosis. Cercus long and thin, truncated and widest at apex, with apophysis at apical third; middle tibia with 5 to 7 long setae, decreasing in length towards base (with two very long subapical setae); middle basitarsus with 5 long setae; fifth and sometimes fourth tarsomeres white haired; femora almost entirely yellow; antenna black; wing vein m-cu strongly sinuate.
Distribution. Congo (Kinshasa), Tanzania, Burundi.
43. Chrysosoma (Chrysosoma) leucopogon (Wiedemann)
Remark. I re-examined material including several specimens determined by C.E. Dyte from the Natural History Museum in London (Grichanov, 1997) and collected in Kenya, Tanzania, on Chagos and Maldives. All males strongly differ from C. leucopogon as described by Bickel (1994a) in hypopygium morphology, belonging apparently to C. snelli described and figured accurately by Curran (1927). Long seta on 7th tergite and unequal arms of cercus figured by Lamb (1922) show that his description of C. leucopogon from Seychelles belongs also to C. snelli. Among other authors, only Parent (1934) recorded this species for western Indian Ocean Region (Mauritius).
Distribution. ?Mauritius, ?Aldabra; Chagos Archipelago, Burma, Sri Lanka, Taiwan, India, Indochina, Java, Sumatra, Thailand, Queensland, New Caledonia, Tahiti.
44. Chrysosoma (Kalocheta) neoliberia Bickel
Type material examined. Paratypes, 3 ??, [red label] / Musee du Congo, Liberia: Du River, Camp N 3 1926, D. J. Bequaert / D. Det. U 1954 / Paratype, Kalocheta liberia Curran.
Diagnosis. Face thinly whitish pollinose, propleural hairs black. Fore coxa with white hairs and black bristles; wing distinctly maculated. Males with hairlike part of arista as long as strap-like part; lower calypter with black hairs.
Distribution. Liberia.
45. Chrysosoma (Chrysosoma) mesotrichum (Bezzi) (Fig. 15)
Type material examined. Holotype, ?, Congo Belg.: P.N.G. Miss H. De Saeger, II/fd/17,9.VII.1952, H. De Saeger, 3763 / Holotypus [red label] / Coll. Mus. Congo (ex coll. IPNCB) / P. Vanschuytbroeck det. 1959, Chrysosoma garambaensis n.sp. [RMCA]. New synonym.
Additional material. 1 ?, Kenya: Kakamega forest, Isecheno stat., pump house trail, 22.VI.1995 (afternoon), N 0º16'46", E 34º46'61", Leg. Earthwatch Team 4; 1 ?, Kenya: Kakamega bridge, Isiukhu river, 17.IX.1995, N 0º16'46", E 34º46'61", Leg. Earthwatch Team 5. 1 ?, Kenya: Kakamega, Virembe nr. trading center, 17.IX.1995, N 0º14'26", E 34º50'93", Leg. Earthwatch Team 5 [NMK]. 1 ?, Congo: Francois / R.I.Sc.N.B. I.G. 24.452. 1 ?, Congo Belge: Kivu, Beni (poste), 18.VI.1953, J. Verbeke, KEA [RINS].
Diagnosis. C. mesotrichum is associated with a group of species having black antenna and numerous long setae on middle tibia and basitarsus. It differs from other species in strongly sinuous vein m-cu, which 1.5 2 times as long as fork-handle M1+2; also in simple cercus with rounded apex and pointed dorsal apophysis, and additional row of short dorsal hairs on first and second joints of middle tarsus, which hardly longer than tarsomere diameter. Cercus with a row of dorsal setae in basal half, with 1 strong and a few fine dorsal setae in apical half, with 2 strong setae and short hairs on apex, which evenly cut and slightly broadened; dorsal dens with 3 or 4 preapical setae. Males with a group of lateral hairs on frons and mostly black femora, whereas females have strong vertical seta and yellow femora.
Distribution. Congo (Kinshasa), Sierra Leone, Uganda, Kenya.
46. Chrysosoma (Chrysosoma) norma Curran
Type material examined. Holotypus, ? [red label] / Musee du Congo, Albertville, Congo, Oct. 1925, Dr. H. Schouteden / Type Chrysosoma norma Curran.
Additional material. 3 ??, Kasai: Terr. de Dekese, Itunda, XI.1959, F.J. Francois / R.I.Sc.N.B. I.G. 24.452.
Diagnosis. Middle basitarsus without long setae; all coxae black, sometimes fore coxa yellow at apex; fore and middle femora black except apical quarter; first three tarsomeres of middle tarsus brownish; lower calypter with black cilia; antenna at least partly yellow-red; cercus with long pointed dorsal apophysis at basal third, thin in the middle, slightly enlarged and setosed at apex, with very thin and sharp subapical dorsal hook.
Distribution. Congo (Kinshasa).
47. Chrysosoma (Chrysosoma) pauperculum Parent
Type material examined. Holotypus, ? [red label] / Musee du Congo, Ituri: Gaduma Malo (Faradji), III.1930, A. Collart / Chrysosoma pauperculum n.sp. Type. O. Parent.
Diagnosis. C. pauperculum is placed within the group of species having no long setae on middle basitarsus. Antenna black. Wing hyaline, vein m-cu sinuate. Only fore coxa yellow. Posterior four coxae black. Middle tibia and tarsus without erect pubescence. Cercus not bifurcated, digitiform, with short dens in the middle.
Distribution. Congo (Kinshasa).
48. Chrysosoma (Chrysosoma) praecipuum Parent
Type material examined. Holotypus, ? [red label] / Musee du Congo, Bambesa, 15.IX.1933, J.V. Leroy / Chrysosoma praecipuum n.sp. Type. O. Parent.
Diagnosis. The species with unclear position within the genus Chrysosoma. It has coxae black, femora mostly black; wing black-brown, whitish along posterior edge, with narrow white transverse band, falling down from R1; lower calypter with pale cilia.
Distribution. Congo (Kinshasa).
49. Chrysosoma (Chrysosoma) pseudorepertum sp.n. (Fig. 16)
Holotype. ?. Congo Belge, Uele: Bili, III.1953 [RINS].
Description. Frons broad, metallic blue-violet, with one fine black lateral seta; postvertical seta positioned as a linear continuation of short postocular setal row. Ocellar tubercle with a pair of strong curved setae and one pair of short hairs. Ventral postcranium covered with dense irregular white hairs. Face metallic blue-green, mostly whitish pollinose; clypeus bulging, separated from eyes; face narrowed, 1.3 times as high as wide under antennae. Palpi and proboscis brown, with pale hairs; palpus also with a pair of black bristles. Antenna mostly orange-brownish, 1.8 times as long as height of head. Scape simple. Pedicel with a ring of strong bristles, at most as long as 1st flagellomere. First flagellomere dark-brown at apex, subtriangular, asymmetric, slightly longer than high. Arista apical, bare and simple. Length ratio of scape to pedicel to first flagellomere to arista, 5 : 5 : 12 : 190.
Mesonotum metallic blue-green; pleura bronze-black, grey pollinose. Two strong posterior and 2 microscopic anterior dorsocentral setae; 3 pairs of strong long acrostichals. Scutellum with two strong bristles, without lateral hairs.
Legs mostly yellow. Fore coxa black except apex; middle and hind coxae and trochanters black; hind femora blackish at extreme apex; hind tibia black at apex; 2nd to 5th segments of fore tarsus brown; hind tarsus entirely black. Fore coxa from the front with numerous white hairs and 3 black subapical setae. Middle coxa with white anterior hairs and 2 black subapical setae; hind coxa with several white external hairs and one black seta in the middle. All femora with light ventral hairs in basal half, slightly longer than femora diameter, gradually decreasing in length from fore to hind femora and from base towards the middle of corresponding femora. Fore tibia with 3 dorsal, 1 anterodorsal and 1 or 2 apical setae. Fore basitarsus with posteroventral row of elongate setulae, at most equal in length to basitarsus diameter. Length ratio of fore coxa to femora to tibia to tarsus (segments from first to fifth), 40 : 78 : 90 : 67 : 17 : 9 : 7 : 8. Middle tibia with 5 or 6 (including apical one) long posterodorsal setae, gradually increasing in length towards apex, and additional posterodorsal row of hairs, slightly longer than tibia diameter, 5 strong anterodorsal, 1 ventral and 3 or 4 apical setae. Middle basitarsus with 4 to 6 long posterodorsal setae and additional posterodorsal row of hairs, twice longer than tarsomere diameter; other tarsomeres without rows or brushes of remarkable hairs. Length ratio of middle coxa to femora to tibia to tarsus (segments from first to fifth), 33 : 100 : 156 : 120 : 30 : 17 : 8 : 8. Hind tibia with 7 dorsal, 4 anterior, 6 short ventral and 3 apical setae. Hind basitarsus with one basoventral seta. Length ratio of hind coxa to femora to tibia to basitarsus (other tarsomeres broken), 21 : 110 : 175 : 85.
Wing widely brownish anteriorly and along veins, with hyaline medial and posterior cells; veins brown. R1 0.4 times as long as wing. M1 convex anteriad. M1+2 and M1 forming obtuse angle. Ratio of part of costa between R2+3 and R4+5 to this between R4+5 and M1, 23 : 7. M2 present as short stub vein and faint curved fold on membrane. Crossvein m-cu strongly sinuate. Ratio of crossvein m-cu, measured along sinuation, to apical part of M 1+2 (fork-handle), 10 : 7. Anal vein faint, anal lobe and alula present. Anal angle acute. Lower calypter brown, with fine, shining light, cilia. Halters black-brown, halter stem yellow at base, thin, 3 or 4 times as long as knob, with short row of black setulae.
Abdomen metallic blue-green, whitish pollinose, with black sutures, short black hairs and long setae. First tergite with pale lateral hairs; sternite with pale and dark hairs. 1st to 6th segments combined twice longer than mesonotum. 7th segment as long as epandrium, with 2 pairs of strong setae; 7th and 8th tergites with long black cilia. Hypopygium small, black. Cercus black, digitiform, with suboval apex and thin long mid-dorsal apophysis, 1/3 as long as cercus; dorsal side of cercus densely covered with long cilia; apex with 2 pairs of thick setae and dense short hairs. Surstylus black, shallow excavated on apex, with short apical setulae and short apicodorsal process.
Female unknown.
Length: body 7.9 mm; antenna 3.5 mm; wing-length 8.7 mm; wing-width 2.2 mm.
Distribution. Congo (Kinshasa).
Diagnosis. C. pseudorepertum is associated with a group of species having yellow antenna and numerous long setae on middle basitarsus and tibia. It differs from C. repertum species in yellow femora and bare 2nd to 5th segments of middle tarsus. It has some similarities with C. katangense, differing from the latter species in long thin pointed mid-dorsal apophysis on cercus and distinctly orange antenna.
50. Chrysosoma (Chrysosoma) schoutedeni Curran (Fig. 17)
Type material examined. Holotypus, ? [red label] / Musee du Congo, Albertville, Congo, Oct. 1925, Dr. H. Schouteden / Holotype Chrysosoma schoutedeni Curran.
Diagnosis. C. schoutedeni is similar to C. mesotrichum and C. consentium, differing mainly in morphology of the hypopygium. Fore and middle femora black in basal third; hind femora entirely black; middle tibia and tarsus except apex yellow. All femora with white ventral hairs in basal half. First and second tarsomeres of middle tarsus with additional row of short dorsal hairs, which are more than twice as long as tarsomere diameter; middle tibia with 7 to 10 long posterodorsal setae; middle basitarsus with 6 to 10 long setae. 2nd to 5th tarsomeres of middle tarsus with black hairs; 5th tarsomere with pile of white hairs. Cercus with long thin dorsal apophysis, longer than width of cercus at apex, with short hairs in apical half of cercus.
Distribution. Congo (Brazzaville), Congo (Kinshasa), Tanzania, Angola, Zambia.
51. Chrysosoma (Chrysosoma) singulare Parent
Type material examined. Holotype ?, Leopoldville, August 1911, Muileman / Chrysosoma singulare n.sp. Type. O. Parent rev., 1933 / Type [red label]. Paratype ?, Mateba, Aug. 1911, Muileman / det. O. Parent 93, Chrysosoma singulare Parent / Paratype [red label].
Additional type material. Holotype, ?, Congo Belg.: P.N.G. Miss H. De Saeger, Akain, 19.V.1950, Rec. H. De Saeger, 529 / Holotypus [red label] / Coll. Mus. Congo (ex coll. IPNCB) / P. Vanschuytbroeck det. 1959, Chrysosoma gracilis n.sp. [RMCA].
Diagnosis. Antenna black. Middle basitarsus without long setae; fore coxa ye
